Nealimyia evenhuisi, Grimaldi, 2016

Grimaldi, David A., 2016, Diverse Orthorrhaphan Flies (Insecta: Diptera: Brachycera) In Amber From The Cretaceous Of Myanmar: Brachycera In Cretaceous Amber, Part Vii David A. Grimaldi, Bulletin of the American Museum of Natural History 2016 (408), pp. 1-132 : 64-66

Nealimyia evenhuisi , new species

Figures 22 , 54

DIAGNOSIS: As for genus, by monotypy.

DESCRIPTION: Based on female. Hunchbacked fly, with large, arched scutum; head attached to thorax just above procoxae, top of head far below top of scutum. Cuticle of fly dark, true color not discernable. Body length 5.05 mm; thorax length 1.95 mm; abdomen length 2.46 mm; wing length 3.96 mm. Head: Slightly hemispherical, with large eyes. Eye (female) occupying all of lateral and most of frontal surfaces of head, not extended to posterior (occipital) region of head. Eye completely bare, no differentiation of facets; posterior margin simple, not emarginate, no transverse line through middle of eye. In lateral view dorsal portion of occiput protruding; ventral portion of eye broader, more rounded than dorsally; eye occupying ventrolateral surface of head (no cheek exposed). Female eyes well separated, lateral margins of frons slightly convergent dorsally, almost parallel; frons bare, no pilosity or pubescence. Ocelli well developed, level with vertex of head (not on mound). Face including clypeus sloped steeply into head, beginning just below antennal bases to deep pit just above pro- boscis. Occiput and postgena partially recessed, but without broad, hemispherical concavity as in Anthracinae . Postgena with median longitudinal suture just below occipital foramen; posterior tentorial pits lateral to suture, just below ventral margin of occipital foramen; foramen apparently not constricted or divided into two; occiput protruding well beyond dorsal margin of eye, setose. Occipital pocket, occipital apodeme apparently absent. Dorsal portion of occiput with median sulcus. Antenna: Insertions (sockets) subcircular, very close, located just above facial depression; antennae held forward. Scape narrow at base, slightly wider distally, slightly longer than pedicel; pedicel slightly broadened distally; flagellum long, slender, drop shaped, unsegmented, tapered to narrow width, with minute apical style at tip (seen in left antenna, tip of right one lost). Mouthparts: Proboscis short, only labellum exposed frontally and laterally; labellum well developed, opened in holotype exposing broad, flat pseudotracheal surface; labellar lobe ear shaped, with ca. 8 pseudotracheae and fringe of fine, stiff setae on lateral rim. Labium very short, barely visible ventrally. Palp very short, not extending beyond labellum, basal palpomere slightly larger than apical one, apex of palp (i.e., pit) not observable. Lacinia and labrum not visible. Thorax: Large, deep, hunched; scutum large, deep, arched; transverse suture well developed. Scutum without macrosetae (bristles), largest setae are fine, short, stiff ones anterior to scutellum; most of scutum with very small, fine setulae; patches of golden, fine, sparse pilosity on supraalar and notopleural regions, portions of katepisternum and anepisternum, and posterior surface of metacoxa. Notopleural suture well developed, completely anteroposterially transverse through thorax. Neck with long antepronotum and lateral cervical sclerites, located just above base of procoxa. Katepisternum relatively small; meron and anepisternum large, well defined; postalar wall dorsally protruding as shallow ledge, ventrally recessed. No proscutellum; scutellum relatively small compared to scutum, dorsal surface entirely with setulae, longer ones on posterior margin; no subscutellum; postnotum short. Wing: Long, slender, W/L 0.29; entirely hyaline, no pterostigma or other infuscation; membrane very faintly and finely wrinkled, with fine microtrichia. Basicosta small, not lappetlike; vein C circumambient, crossvein h heavily sclerotized; Sc 0.48× length of wing, complete; R 1 very close to and parallel to Sc, almost fused apically; faint sc-r crossvein present. R 2+3 0.78× wing length, tip upturned; R 4 -R 5 forked, tips of fork encompassing wing tip, R 4 slightly sinuous and longer than R 5. Cell d large, r-m near apex of cell; two branches of M present (M 1, M 2); CuA 1 arising preapically from cell d. Cell br much longer (1.56×) than cell bm; stem of M heavily sclerotized. CuA 2 and A 1 meeting wing margin separately (cell cup open); anal lobe and alula narrow; upper calypter very small. Legs: Slender. Procoxa ca. 2× length of meso- and metacoxae. Femora with short, decumbent setae only. Lengths of femora: metafemur> meso> profemur. Tibia slightly longer than respective femur; protibia without macrosetae; mesotibia with dorsolongitudinal row of eight stiff, short setae; metatibia with dorsolongitudinal and ventrolongitudinal row of 8–6 setae, respectively. Tibial spurs 0-1-0; mesotibial spur slightly longer than apical width of tibia. Basitarsus longer than combined distal tarsomeres. Claws well developed, simple; empodium setiform; pulvilli are short pads, shorter than claws. Abdomen (female): Significantly more slender than thorax in dorsal view, slightly longer than thorax. Tergite 1 simple, not divided; tergites 2–7 little differentiated in size, all with dense vestiture of small, decumbent setae. Terminalia (female): Posterior margin of tergite 7 and sternite 7 (possibly t8 + s8, which are recessed within 7) with long, dense brush of fine, golden setae, forming a thick “basket.” Presence/absence of acanthophorite spines not observable.

TYPE: Holotype, female, AMNH Bu-SE2/7.

ETYMOLOGY: Patronym for Neal Evenhuis, dipterist, bombyliid specialist, colleague.

COMMENTS: The fly lies in a slabbed portion of a runnel, 13 × 9 × 4 mm, which also contains some internal flow surfaces and a light suspension of very fine bubbles and particles. The fly is complete and very well preserved.

Figures

FIG. 22. Nealimyia evenhuisi (Bombyliidae), new genus and species, holotype, AMNH Bu-SE2/7. A. Head and thorax, right lateral view, with detail of antenna. B. Wing. C. Detail of head, oblique ventral view of anterior portion. D. Detail of head, frontal view (only antennal sockets shown).

FIG. 54. Nealimyia evenhuisi, new genus and species (Bombyliidae), holotype, AMNH Bu-SE2/7. A. Habitus, right lateral. B. Head and thorax, right lateral. C. Head, right lateral. D. Terminalia, right lateral, showing dense brush of fine setae.