Normyia woodleyi, Grimaldi, 2016

Grimaldi, David A., 2016, Diverse Orthorrhaphan Flies (Insecta: Diptera: Brachycera) In Amber From The Cretaceous Of Myanmar: Brachycera In Cretaceous Amber, Part Vii David A. Grimaldi, Bulletin of the American Museum of Natural History 2016 (408), pp. 1-132 : 21-23

Normyia woodleyi , new species

Figures 7 , 37–39

DIAGNOSIS: Based on male and female. Basal and apical flagellomeres (1, 8) short, <2× lengths of adjacent flagellomeres (2, 7); R 5 turned slightly upward ; cell d large, L/ W 2.14 (vs. 1.75–2.0); mesotibia with 1 stout spur; terminal abdominal segments of female not telescoping.

DESCRIPTION: Male holotype, paratype. Body length 2.31, 2.56 mm (respectively), thorax length 0.98, 1.01 mm; wing length 1.59, 2.02 mm; abdomen length 0.93, 1.20 mm. Female paratype: body length 2.85, thorax length 1.01 mm, wing length 1.94 mm, abdomen length 1.48 mm. Head: Large, significantly broader than thorax, approximately hemispherical in shape with occiput dark, flat to slightly recessed, with virtually no setae. Ocelli on raised tubercle, with anterior ocellus directed forward, posterior ocelli directed laterally. Eye: Male: Large, holoptic, occupying nearly all of nonoccipital portion of head; facets with dorsoventral differentiation, dorsal ones ca. 2× diameter of ventral ones, a discrete transverse separation between the two. Eyes completely bare; medial/inner margins meeting just anterior to ocellar triangle, strongly diverging just above antennae, slightly emarginate near antennal bases, inner margins ventrally slightly divergent posteriad; essentially no cheek exposed. Female: Frons widely separated, lateral margins parallel. Antenna: With 8 flagellomeres, submoniliform, tapered in width apically; flagellomeres 1 and 8 longest, flagellomere 1 width 5× that of flagellomere 8; no apical stylus; all flagellomeres with sensillar foveae; scape funnel shaped, pedicel flask shaped, with apical whorl of setae, length of antenna 0.35 mm; antennal sockets very close, possibly touching medially. Mouthparts: Clypeus and palps small, recessed into shallow concavity of face, face is fully ventral; clypeus small, narrow, triangular; labrum short; palps small, light, 2-segmented, basal palpomere slightly longer and narrower than apical one, palp overall slightly shorter than clypeus. Labellum well developed, fleshy, exposed well beyond oral margin. Thorax: Relatively short, deep in lateral view, dark (notum probably somewhat metallic, given tiny blue-green reflections from cuticle). Scutum and scutellum with very fine, setigerous punctation; setulae very fine, short. Antepronotum very well developed; postpronotum large, joined to lateral margins of antepronotum; postpronotal lobe narrow, triangular; anterior third of scutum with pair of very faint paramedian sulci. Transverse suture very well developed, deeply impressed, inner apices converging posteromedially; suture forming rounded lobelike portion of scutum posteriorly. Small proscutellum present; scutellum narrow, length slightly less than width, no spines. Posterior portion of anepisternum large, bulging; katepisternum ventrally extended, nearly pendulous, with ventral margin nearly at same level as ventral margin of mesocoxa; medial margins of both katepisterna meeting ventrally. Wing: W/L 0.37. Sc complete, long, 0.5× wing length; C ends slightly past tip of R 5. R and R 1 close and parallel to Sc, diverging slightly at apices. Rs very short, 0.25× length of R 1; R 2+3 slightly shorter than R 1, convergent, apices of veins nearly touching; fork of R 4 -R 5 long, base of fork only slightly distal to level of cell d apex; R 4 sinuous, R 5 straight, tip slightly upturned, ending before wing tip. Faint pterostigma between Sc, R 1, and R 2+3, r-m crossvein at middle of cell d; base of M between cells br and bm very faint, not tubular or at all sclerotized. M 1 faintly curved; apices of M 1, M 2, CuA 1 tapered and evanescent, not meeting wing mar- gin; bases of M 1 and M 2 either meeting truncate apex of cell d separately (e.g., left wing of holotype), or cell d apex pointed and joined to bases of R 4 and R 5 into an “X” (i.e., right wing of holotype). Cell d small, W/L 0.43; cell br longer than bm, both cells narrow. CuA 2 arched, meeting A 1 well before wing margin. Anal lobe slender; A 2 short and extremely faint; alula small. Legs: Relatively short, covered with fine setulae, tibial spurs 0-1-0, length of mesotibial spur slightly shorter than width of tibial apex, distal half of spur heavily sclerotized. Length of femur> tibia> basitarsomere = combined lengths of tarsomeres 2–5. Empodium pulvilliform; empodium and pulvilli small, significantly shorter than pretarsal claws. Abdomen: Broad, relatively short, moderately flattened dorsoventrally. Tergites 1–5 each with transverse furrow impressed near posterior margins, furrows deepest on tergites 2–4, furrow lengths ca. 0.8× tergite width; 7 tergites exposed (7th very small) in male, 10 in female. Sternites 1–7 well developed (if an 8th, then not visible). Male Terminalia: Well developed, exposed. Gonocoxa simple, short, cylindrical, fully articulating with gonostylus; gonostylus somewhat scoop shaped, broad in lateral view, with slightly hooked apical spine; smaller preapical median spine. Proctiger small, approximately triangular (apex narrow), cercus 1-segmented, ovoid, lateral to epiproct. Female Terminalia: Cercus 2-segmented, segments approximately equal in length, basal cercomere slightly thicker; terminalia not telescoping. Sternite 10 possibly present.

TYPES: Holotype, male, AMNH Bu-SE04B . Paratype male, AMNH Bu-SE04A ; paratype female, JZC Bu-001.

ETYMOLOGY: Patronym, for dipterist Norman Woodley.

COMMENTS: The holotype and paratype, AMNH Bu-SE4A, B are very well-preserved males from the same piece, originally a cabochen 12 × 8 × 7 mm, which are now separated into small polished cubes each 6 × 5 × 3 mm. The paratype is partial (lost left side of thorax and head), but the internal clearing better resolves some sutures. The male holotype is complete and superbly preserved ; there is a mycetophilid syninclusion.

Figures

FIG. 7. Normyia woodleyi, new genus and species (Stratiomyidae). A–F: Holotype male, AMNH Bu-SE04b. G: Paratype female, JZC Bu-001. A. Wing. B. Male terminalia, oblique dorsal view. C. Male terminalia, oblique ventral view. D. Palp, holotype (above), female paratype (below). E. Mesotibial apex with spur. F. Antenna. G. Female terminalia, posterolateral view. Panels C−E to same scale.

FIG. 37. Normyia woodleyi, new genus and species (Stratiomyidae), holotype male AMNH Bu-SE004b. A. Habitus, dorsal. B. Head, oblique dorsal view. C. Terminalia, posteroventral view.

FIG. 38. Normyia woodleyi, paratype male AMNH Bu-SE004a. A. Habitus, right lateral view. B. Detail of abdomen, showing deeply impressed tergal sulci. C. Right wing.

FIG. 39. Normyia woodleyi, paratype female JZC Bu-001. A. Habitus, right lateral. B. Habitus, left lateral. C. Habitus, dorsal. D. Head, right lateral. E. Terminalia, oblique left lateral.