Ainigmapsychops inexspectatus, Makarkin & Archibald, 2014

Makarkin, Vladimir N. & Archibald, S. Bruce, 2014, An unusual new fossil genus probably belonging to the Psychopsidae (Neuroptera) from the Eocene Okanagan Highlands, western North America, Zootaxa 3838 (3), pp. 385-391 : 387

Ainigmapsychops inexspectatus sp. nov.

( Fig. 1 )

Description. Forewing 7.2 mm long as preserved (estimated complete length ca. 15 mm), 6.5 mm wide. Costal space very broad medially and probably basally, strongly narrowed toward apex. Most preserved subcostal veinlets deeply forked, others simple. No costal crossveins. ScP stout, especially proximally. Subcostal space moderately narrow, with weakly-developed crossveins (six detected, but possibly more present). RA space narrow, only slightly wider than subcostal space, with crossveins irregularly spaced (ten detected, possibly more present). RP nearly smooth, with 15 preserved branches. RP2, RP3 fused at level of termination of AA1. Preserved branches of RP not forked before outer gradate series. Radial crossveins numerous, mostly arranged in four gradate series, but proximal crossveins mainly spaced irregularly; first gradate series running close to RP stem; second series irregular, short, located distad proximal crossveins; third (intermediate) series strongly irregular, occupying central wing area; fourth (outer) series regular. Fork of M not preserved. MA incomplete, probably fused with RP1 at outer gradate series (or terminating on RP1). MP deeply forked at outer gradate series. Intramedial crossveins rare, two detected. Mediocubital space (between MP, CuA) broad, with many irregularly spaced crossveins (all curved or oblique; two crossveins connected by additional crossvein). CuA markedly convex, quite deeply forked distally, pectinately branched with six branches (four rather deeply forked, two simple). CuP running parallel to hind margin, deeply forked distally, pectinately branched with two simple branches. Mediocubital, intracubital and cubitoanal spaces with many irregularly spaced crossveins (all curved or oblique; four pairs of crossveins connected by additional crossveins). AA1 long, running parallel to hind margin, pectinately branched with four preserved branches (three deeply forked, one simple, one incomplete). No crossveins between branches of AA1. Wing membrane appears more or less one-colored, dark, here preserved as brownish.

Material. Holotype SRUI 99-96 - 76 (part only), collected by Gregg Wilson on April 28, 2013. A rather wellpreserved central part of a forewing, housed in the collection of the Stonerose Interpretive Center , Republic, Washington, U.S.A.

Type locality and horizon. Tom Thumb Tuff Member of the Klondike Mountain Formation , exposure A0307, Republic, Washington, U.S.A.; early Eocene (Ypresian) .

Etymology. From the Latin inexspectatus [- a, - um], unexpected, in reference to the surprise of finding this odd wing.

Figures

FIGURE 1. Ainigmapsychops inexspectatus gen et sp. nov., holotype SRUI 99-96-76. A, specimen as preserved; B, forewing venation. Scale bar = 2 mm.