Cuccodorodes koshiensis, Yin, 2018

Yin, Zi-Wei, 2018, Cuccodorodes gen. nov., an apterous Batrisitae from the tropical Himalayas (Coleoptera: Staphylinidae: Pselaphinae), Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2), pp. 321-330 : 328

Cuccodorodes koshiensis sp. nov.

( Figs 2A , 3 C–D, 5 E, 6 D–F, 7 )

Type material (14 ♂♂ 33 ♀♀). HOLOTYPE: NEPAL: ♂, ‘E. NEPAL: KOSI, Val. Induwa Kola, 2800 m, 15.iv. (19)84, Löbl – Smetana’ ( MHNG). PARATYPES: NEPAL: 13 ♂♂ 30 ♀♀, same label data as holotype; same collector, also from KOSI; 1 ♀, ‘ Goru Dzure Dara , 3600 m, 9.iv.84’; 1 ♀, ‘ Goru Dzure Dara , 3350 m, 9.iv.84’; 1 ♀, ‘ Crete , S. Mangsingma , 2800 m, 7.iv.84’ (4 exs in SNUC, rest in MHNG).

Description. Male. General habitus stout ( Fig. 2A ), body reddish brown, pronotum slightly darker, mouthparts and tarsomeres slighter in color, BL 3.02–3.36 mm. Head about as long as wide, HL 0.56–0.58 mm, HW 0.57–0.59 mm; vertexal foveae located far below level of posterior margin of eyes, sulcus evenly convergent anteriorly, more or less V-shaped ( Fig. 3C ), median carina extending anteriorly from near head base to meet sulcus; eyes small, slightly protruding laterally, each eye composed of about 10 facets; AnL 1.18–1.20 mm. Pronotum wider than long, PL 0.58–0.62 mm, PW 0.68–0.69 mm; median longitudinal sulcus extending from small median antebasal fovea to near apex, discal longitudinal sulci slightly curved, lateral longitudinal sulci strongly sinuate, broadening at basal half and densely setose. Elytra much wider than long, EL 0.72–0.82 mm, EW 0.87–0.93 mm. Abdomen widest at tergite 1 (IV), AL 1.13–1.16 mm, AW 0.89–1.00 mm; sternite 1 (IV) with broad and moderately thickened arched median projection ( Fig. 3D ), surface of projection with scaly microsculpture, area surrounded by projection broadly triangular; median area of sternite 3 (V) lacking microsculpture; sternite 7 (IX) with membranous basal half broadly expanded ( Fig. 6D ). Length of aedeagus ( Figs 6 E–F) 0.64 mm, ventral lobe strongly elongate, protruding posteroventrally; two markedly elongate sclerites at apex of ventral lobe and left side.

Female. General habitus similar to male; with slightly shorter antennae; each eye composed of about 9–10 facets; mesotibia simple; none of available specimens has median projection on tergites 3–4 (VI–VII); sternite 2 (IV) unmodified; genital complex ( Fig. 5E ) with basolateral extensions roundly narrowing posteriorly. Measurements: BL 2.93–3.02 mm, AnL 1.13 mm, HL 0.56–0.58 mm, HW 0.56–0.58 mm, PL 0.58 mm, PW 0.68 mm, EL 0.71–0.74 mm, EW 0.89–0.92 mm, AL 1.10–1.12 mm, AW 0.96–0.97 mm.

Etymology. The new species is named after Koshi Zone, where the type locality lies; adjective.

Distribution. Nepal: Koshi (= Kosi) Zone ( Fig. 7 ).

Figures

Fig. 2. Dorsal habitus of Cuccodorodes species. A – C. koshiensis sp. nov., male; B – C. weiperti sp. nov., male.

Fig. 3. Male head and pronotum (A, C, E), and sternites 2–3 (IV–V) (B, D, F) of Cuccodorodes darjeelingensis sp. nov. (A–B), C. koshiensis sp. nov. (C–D), and C. weiperti sp. nov. (E–F).

Fig. 5. Morphology of Cuccodorodes gen. nov., based on C. koshiensis sp. nov.unless specified otherwise.A – prothorax, in dorso-lateral view; B – mesoand metaventrite; C – internal foveal structure of tergite 1 (IV) and sternite 2 (IV); D–E – female genital complex, in dorsal view (D – C. darjeelingensis sp. nov.; E – C. koshiensis sp. nov.). Abbreviations: blf – basolateral foveae; dls – discal longitudinal sulcus; laf – lateral antebasal foveae; lls – lateral longitudinal sulcus; lmcf – lateral mesocoxal foveae; lmsf – lateral mesoventral foveae; lmtf – lateral metaventral foveae; mbf – mediobasal foveae; mls – median longitudinal sulcus; mmsf – median mesoventral foveae; mp – median projection; p – pit. Scale bars: 0.2 mm.

Fig. 6. Male sternite 7 (IX) (A, D, G), and aedeagus (B–C, E–F, H–I) of Cuccodorodes darjeelingensis sp. nov. (A–C), C. koshiensis sp. nov. (D–F), and C. weiperti sp. nov. (G–I). Scale bars: 0.1 mm in A, D, G; 0.2 mm in B–C, E–F, H–I.

Fig.7. Map showing the distribution of Cuccodorodes species.