Micrathyria Kirby, 1889

Mauffray, William F. & Tennessen, Kenneth J., 2019, A Catalogue and Historical Study of the Odonata of Ecuador, Zootaxa 4628 (1), pp. 1-265 : 177-182

publication ID

https://doi.org/10.11646/zootaxa.4628.1.1

publication LSID

lsid:zoobank.org:pub:E4C2F52F-9A1A-4C55-B03B-C87B2B4C4676

DOI

https://doi.org/10.5281/zenodo.5921552

persistent identifier

https://treatment.plazi.org/id/D076D419-B5A3-FF63-FF23-F760201EF8CD

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Plazi

scientific name

Micrathyria Kirby, 1889
status

 

Micrathyria Kirby, 1889

A large group of small to medium-sized dragonflies ( Fig. 102 View FIGURE 102 ) in need of revision. Heckman (2006) devised a key to species based on the scattered literature. Twenty-one species have been recorded for Ecuador, plus there are a number of specimens with questionable identity that could increase this number. The group occupies a wide variety of habitats including vegetated lakes, ponds, and sloughs, both open and shaded.

Micrathyria aequalis ( Hagen, 1861) . Publ. Recs. ECU: Muttkowski (1910 —as M. septima and proposed synonymy with M. aequalis ), Paulson (1983), Tsuda (1986, 1991), Silsby (2001), (H06), (P11), (H14); GY: * Calvert (1906), Campos (1922). Also CEN, COL, GUF, GUY, SUR, VEN: (H06), (P11); plus TTO: (P11).

Addl. Data. EO, Arenillas: 1 ♂, farm pond, seeps, 11 km S of Arenillas, on road to Alamar (-3.6436, - 80.0989, 95m), 10 iv 2008, leg. & det. JTJ [JTJ]; 2 ♂♂, same data but leg. & det. KJT [KJT]; Machala: 1 ♀, borrow pit pond, Hwy. 25, S of Machala (-3.4750, -79.9778), 10 iv 2008, leg. & det. KJT [KJT]; Santa Rosa: 1 ♂, pond, 12 km E of Arenillas, Hwy. 25 (-3.5244, -79.9831, 30m), 29 iii 2008, leg. & det. KJT [KJT]; 3 ♂♂, same data but leg. & det. JTJ [JTJ]; 2 ♂♂, same data but 9 iv 2008, leg. & det. KJT [KJT]; Piñas: 1 ♀, Piedras (-3.63, -79.92, 300m), 6 vii 1941, leg. D. Laddey, det. KJT [FSCA]; GY, Daule: 1 ♂, Nobol, 5.5 km N, (30m), 12 i 1978, leg. P. Spangler & J. Anderson, det.? [USNM]; Guayaquil: 1 ♂, Guayaquil, leg. F. Campos, det. MJW [ANSP]; LR, Babahoyo: adults, 20 km N of Babahoyo, 22 vi 1975, leg. J. Cohen, A. Langley & P. Monnig, det.? [USNM]; Ventanas: 1 ♂, pond/stream 7.2 km E of Hwy. E25, 9 km SW of Quinzaloma (-1.2539, -79.3703, 100m), 21 ii 2010, leg. & det. JTJ [JTJ]; MN, El Carmen: 1 ♂, El Carmen, i 1983, leg. & det.? [QZAC]; 1 ♂, borrow pits about 18 km NW of El Carmen (-0.1350, -79.5797, 185m), 2 iii 2010, leg. & det. JTJ [JTJ].

* Micrathyria artemis Ris, 1911 . Publ. Recs. ARG, BRA, GUF, GUY, PER, SUR, VEN: (H06), (P16).

Addl. Data. NA, Tena: 1 ♂, Puerto Napo, 3.2 km E on Jatun Sacha Road, 23 vii 1996, leg. & det. SWD [FSCA]; SU, Shushufindi: 1 ♂, Limoncocha, lake edge (-0.4000, -76.6167, 269m), 8 ii 1971, leg. D. Pearson, det. DRP [ DRP]; 1 ♂, same data but 26 vii 1977, leg. DRP & S. Hills, det. DRP, checked by RWG [DRP].

Micrathyria atra ( Martin,1897) . Publ. Recs. ECU: (H06), (P11), (H14). Also ARG, BRA CEN, GUF, GUY, PER, SUR, TTO, VEN: (H06), (P11).

Addl. Data. MS, Santiago: 1 ♂, [Patuca], [about 1 mi N], 43.4 km S of Sucua, large pond, [about 5 km N of Mendez road jct] (-2.75, -78.27, 488m), 7 xi 1997, leg. & det. KJT [KJT]; 1 ♀, same data but leg. & det. WFM [FSCA]; 3 ♂♂, same data but 8 xi 1997 [FSCA, LSAM, QZAC]; 7 ♂♂, 2 ♀♀, same data but leg. & det. JJD [JJD]; 2 ♂♂, 1 ♀, same data but leg. & det. KJT [KJT]; NA, Tena: 1 ♂, forest pool 14.8 km E of Puerto Napo on J.S.B.S. Road, 15 vi 1995, leg. & det. SWD [JJD]; 1 ♂, same data but leg. & det. KJT [KJT]; 1 ♂, Río Sinde, 5.3 km E of Puerto Napo (-1.0500, -77.7481, 450m), 15 vi 1995, leg. & det. KJT [KJT]; 1 cop. pair, Puerto Napo, 14.8 km E on J.S.B.S. Road, forest pool, 15 vi 1995, leg. & det. SWD [SWD]; 2 ♂♂, 1 ♀, J.S.B.S., Pond #14, 18–23 xi 1991, leg. E. McCallie, det.? [USNM]; OR, Loreto: 2 ♂♂, S of Loreto, forest pond, (-0.8141, - 77.3371, 350m), 2 xi 2013, leg. & det. CDK [ FSCA]; 1 ♂, N of Loreto, pond, (-0.8205, -77.3202), 2 xi 2013, leg. & det. KJT [ FSCA]; SU, Cascales: 1 ♂, pool, 58 km W of Lago Agrio, 20 viii 1980, leg. & det. SWD [ SWD]; Shushufindi: 26 ♂♂, 2 ♀♀, 1 cop. pair, Limoncocha   GoogleMaps , Río   GoogleMaps Napo (-0.4000, -76.6167, 269m), 2–26 xi 1980, leg. & det. MJW [ FSCA]; 3 ♂♂, 2 ♀♀, same data but palm swamp, 25 vii 1977, leg. & det. DRP [ DRP].

Micrathyria caerulistyla Donnelly, 1992 . Publ. Recs. ECU: (H06), (P11), (H14). Also CEN: (H06), (P11).

Addl. Data. MS, Santiago: pond along road between Sucua and Mendez (-2.6833, -78.2283, 600m), 6 xi 1997, leg. & det. TWD [SWD]; NA, Tena: 6 ♂♂, Puerto Napo, 3.2 km E on J.S.B.S. Road, ponds and flowing marsh (-1.0456, -77.7467, 420m), 23 vii 1996, leg. & det. WFM [FSCA, JJD, LSAM, QZAC, USNM]; 4 ♂♂, same data but leg. & det. KJT [KJT]; 4 ♂♂, same data but leg. & det. SWD [RWG, SWD]; 3 ♂♂, 5.3 km E of Hwy. 45 on J.S.B.S. Road, Río Sinde (-1.0500, -77.7481, 450m), 16 x 1988, leg. & det. SWD [FSCA, SWD]; 1 ♂, same data but 9 x 2009, leg. & det. JTJ [JTJ]; OR, Francisco de Orellana: 5 ♂♂, Puerto Francisco de Orellana, 1.7 km W on Loreto Road from Coca-Sachas Road, stream, pools & flowing shallow marsh (-0.4367, -77.0017, 258m), 13 vi 1995, leg. & det. WFM [ FSCA, QZAC, USNM]; 1 ♂, 1 cop. pair, same data but leg. & det. KJT [ KJT]; 2 ♂♂, same data but leg. & det. SWD [ SWD]; 1 ♂, same data but 18 xi 1997, leg. & det. WFM [ FSCA]; 1 ♂, same data but leg. & det. TWD [ SWD]; 2 ♂♂, [ FSCA, JJD]; 1 ♂, 12.3 km W of Coca Road on Loreto Road (820), 13 vi 1995, leg. & det. SWD [ SWD]; 1 ♂, P.N.Y., 6.2 km W of Yasuni Road on PC 29-30, pool/stream, 10 vi 1995, leg. & det. SWD [ SWD]; 1 ♀, [Pompeya Sur] S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto   GoogleMaps (-0.60, -76.47, 229m), 17 vii 1996, leg. & det. SWD [ SWD]; 1 ♂, P.N.Y., 6.3 km S of Yasuni Road on Waorani Reserve Road, marshy pond (-0.7120, -76.4745, 259m), 10 vi 1995, leg. & det. WFM [ FSCA]; 1 ♂, P.N.Y., 2.1 km E of Yasuni Biological Research Station pond and swamp (-0.6741, -76.3685, 244m), 15 xi 1997, leg. & det. WFM [ FSCA]; SU, Shushufindi, 1 ♂, Limoncocha, lakeshore & nearby areas (-0.4000, - 76.6167, 269m), 4 ix 1978, leg. KWK, det. MJW [ FSCA]; La Selva Lodge, Lake Mandicocha   GoogleMaps & outlet stream (-0.4983, -76.3733, 200m), 25 xi 1997, leg. & det. TWD [ SWD]; ZC, Yantzaza: 1 ♂, slow stream 7 km N of El Pangui, 6 xi 1997, leg. & det. KJT [ KJT].

Micrathyria catenata Calvert, 1909 . Publ. Recs. ECU: (H06), (P11), (H14); ES: Daigle et al. (1997); NA: Tennessen (1996b); SU: * Westfall (1992). Also ARG, BRA, CEN, GUF, PER, SUR, TTO, VEN: (H06), (P11); plus COL: Palacino et al. (2017).

Addl. Data. ES: San Lorenzo: [FSCA, JJD, KJT]; MS, Santiago: [FSCA, JJD, KJT, LSAM, QZAC, TWD, USNM]; Taisha: [DRP]; NA, Archidona: [FSCA]; Carlos Julio Arosemena Tola: [FSCA]; Chaco: (KWN), Tena: [DRP, FSCA, JJD, KJT, SWD, USNM]; OR, Francisco de Orellana: [FSCA, KWK, KJT, LSAM, QZAC, RWG, SWD, TWD, USNM]; SU, Cascales: [SWD]; Shushufindi: [FSCA, KJT]; PA, Pastaza: [FSCA, KJT, QZAC, USNM].

Micrathyria dictynna Ris, 1911 . Publ. Recs. ECU: (H06), (P11), (H14); MS: Tennessen (2006); OR: von Ellenrieder & Garrison (2011); SU * Donnelly (1998). Also CEN, VEN: (H06), (P11); plus COL: (P&P).

Addl. Data. MS, Huamboya: 2 ♂♂, small stream w/ pools, forested, near Río Pasanac [57 km N of Macas], Hwy. 45, N of river, E side of Hwy. (-1.9550, -77.8567), 16 ix 2005, leg. & det. JJD [JJD]; 1 ♂, 1 ♀, same data but leg. & det. KJT [KJT]; NA, Tena: 2 ♂♂, J.S.B.S., pond #13, 23 xi 1991, leg. E. McCallie, det.? [USNM]; 1 ♂, Río Sinde, pond, 5.3 km E of Puerto Napo (-1.0517, -77.7544, 450m), 13 vii 1996, leg. & det. KJT [KJT]; 1 ♂, Río Chinchipino, oxbow pool, 27 km E of Puerto Napo (-1.0728, -77.5864, 385m), 13 xi 1997, leg. & det. KJT [KJT]; OR, Francisco de Orellana: 1 ♀, Yuturi , 2 iii 1990, leg. & det.? [ QZAC]; 2 ♂♂, T. B.S., USFQ, Guaca- mayo trail between markers 450–2100 (-0.6350, -76.1533, 237m), 2009, leg. & det. RWG & NVE [ NVE, RWG]; Loreto: 1 ♂, forest about 12 km NE of Loreto (-0.6217, -77.2600, 365m), 15 x 2009, leg. KJT [ KJT]; same data but 1 ♂, 3 xi 2013, leg. & det. KJT [ FSCA]; 1 ♂, S of Loreto, forest pond, (-0.8141 -77.3371, 350m), 2 xi 2013, leg. & det. CDK [ FSCA]; PA, Mera: 1 ♂, seepage stream, Los Copales Cabanas between Mera and Shell, W of Puyo, Hwy.   GoogleMaps 50, 22 ix 2005, leg. & det. JJD [ JJD]; SU, Lago Agrio: 1 ♂, swamp, 17 km S of Lago Agrio, 19 viii 1980, leg. & det. SWD [ SWD]; Shushufindi: 1 ♀, Limoncocha, Río   GoogleMaps Napo (-0.4000, -76.6167, 269m), 11 ix 1973, leg. B. Drummond III, det.? [ FSCA]; same data but forest edge, 25 vii 1977, leg. & det. DRP [ DRP]; 1 ♂, stream about 3 km NW of Shushufindi (-0.1664, -76.6917, 290m), 13 v 2009, leg. & det. KJT [ KJT]; 1 ♂, La Selva Lodge; forest (-0.4983, -76.3733, 200m), 22 xi 1997, leg. & det. TWD [ SWD]; 1 ♀, La Selva, Manicocha, 12 x 1988, leg. & det. SWD [ SWD]; 1 ♂, 1 ♀, La Selva, Garzacocha, 12 x 1988, leg. & det. SWD [ SWD].

Micrathyria didyma (Selys in Sagra, 1857) . Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), Steinmann (1997b as M. didyma didyma ), Silsby (2001), (H06), (P11), (H14); EO: Campos (1927); GY: * Calvert (1906), Ris (1911a), Campos (1922); IM: Ris (1911a—as M. didyma didyma ), Campos (1922). Also CEN, COL, GUF, GUY, SUR, VEN: (H06), (P11); plus BRA: (H06); plus TTO: (P11).

Addl. Data. GY, Guayaquil: 1 ♂, Guayaquil (-2.20, -79.90, 8m), date?, leg. F. Campos, det. P. Calvert, checked WFM [FSCA].

Micrathyria hippolyte Ris, 1911 . Publ. Recs. ECU: (H06), (P11), (H14); NA: * Tennessen (1996b). Also BRA, GUF, GUY, PER, SUR, VEN: (H06), (P11).

Addl. Data. NA, Tena: 1 ♂, [Puerto Napo], [upstream near jct with Río Mulatos] Jatun Yacu Río Napo watershed (-1.05, -77.98, 700m), 25 ii 1937, leg. WCM, det.? [ FSCA]; 2 ♂♂, pond, road to J.S. B.S., 14.8 km E of Puerto Napo, 15 vi 1995, leg. & det. KJT [ KJT]; 2 ♂♂, 1 ♀, same data but leg. & det. SWD [ SWD]; SU, Shushufindi: 1 ♂, Limoncocha   GoogleMaps (-0.4000, -76.6176, 269m), 28 viii 1980, leg. & det. SWD [ SWD]; 1 ♀, same data but forest trail, 27 ix 1971, leg. D. Pearson, det. DRP [ DRP]; 1 ♀, same data but 24 i 1972 [ DRP]; 1 ♂, same data but 19 ii 1972 [ FSCA]; 1 ♂, same data but forest stream, 9 iii 1972 [ DRP]; 1 ♂, same data but forest edge, 24 vii 1977, leg. & det. DRP [ DRP].

Micrathyria iheringi Santos, 1946 . Publ. Recs. ECU: * Daigle et al. (1996), (H06), (P11), (H14). Also BRA: (H06), (P11).

Addl. Data. MS, Huamboya: 1 ♂, Macas, 58 km N on Hwy. 45 (70 km N of Sucua), roadside ditch and tribs. near Río Pasanac   GoogleMaps (-1.9447, -77.8536, 762m), 10 xi 1997, leg. & det. WFM [ FSCA]; 1 ♂, (labeled “PA”), small stream about 1 km S of Río Pasanac   GoogleMaps (-1.9503, -77.8631, 820m), 16 ix 2005, leg. & det. KJT [ FSCA]; 1 ♂, same data but 16 ix 2005 [ KJT]; NA, Tena : 1 ♂, Tena (-0.98, -77.8, 500m), date?, leg. F. Campos, det. KJT [ FSCA]; 1 ♂, J.S. B.S., 18 x 1988, leg. & det. SWD [ SWD]; 1 ♂, same data but Pond #8, 25 xi 1991, leg. E. McCallie, det.? [ USNM]; 2 ♂♂, Río Sinde   GoogleMaps , slough; 5.3 km E Puerto Napo (-1.0500, -77.7481, 450m), 16 x 1988, leg. & det. SWD [ RWG, SWD]; 1 ♂, same data but 13 vii 1996, leg. & det. SWD [ SWD]; 4 ♂♂, pool near Río   GoogleMaps Napo, 3.2 km E of Puerto Napo (-1.0456, -77.7467, 420m), 23 vii 1996, leg. & det. KJT [ KJT]; 1 ♂, same data but leg. & det. SWD [ SWD]; 1 ♂, Puerto Napo, 17.2 km E on J.S.B.S. Road, pond near stream (340m), 22 vii 1996, leg. & det. SWD [ SWD]; 1 ♂, Puerto Napo, 2 nd stream E of Río Sinde on J.S.B.S. Road, 23 vii 1996, leg. & det. SWD [ SWD]; OR, Francisco de Orellana: 1 ♂, Comunidad El Rocio, E on Rt   GoogleMaps E20, pond, (-0.5056, -77.1566), 27 x 2013, leg. & det. KJT [ FSCA]; Loreto: 3 ♂♂, Pano , 12 ix 1990, leg. OSF, det.? [USNM]; SU, Shushufindi : 1 ♂, trails, small streams 3 km W of Shushufindi, 2+ km N off Hwy   GoogleMaps . 26 at km marker 17 (-0.1664, -76.6917, 290m), 13 x 2009, leg. & det. JTJ [ JTJ]; ZC, Zamora : 3 ♂♂, Zamora (-4.07, -78.95, 1000m), 21 x 1941, leg. D. Laddey, det. JTJ [ FSCA].

Micrathyria laevigata Calvert, 1909 . Publ. Recs. ECU: Ris (1911—as M. didyma laevigata ), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997b—as M. didyma laevigata ), (H06), (P11), (H14); SD: Daigle et al. (1997). Also CEN, COL, PER, VEN, TTO, BRA: (H06), (P11).

Addl. Data. OR, Francisco de Orellana: 1 ♂, [La Nueva Esperanza] [across Río Napo from] near Primavera “Limoncocha” Río   GoogleMaps Napo & Lake Taracoa   GoogleMaps (-0.45, -76.77, 300m), 4 xi 1980, leg. & det. MJW [ FSCA]; La Joya de los Sachas: 2 ♂♂, 2 ♀♀, Pompeya. 5 km W of Puerto Pompeya Ferry Landing, roadside ditches (-0.4444, - 76.6263, 238m), 19 vii 1996, leg. & det. WFM [ FSCA, LSAM, QZAC]; 1 ♂, same data but leg. & det. JJD [ JJD]; 3 ♂♂, same data but leg. & det. KJT [ KJT]; SU, Shushufindi: 3 ♂♂, Limoncocha, Río   GoogleMaps Napo (-0.4000, -76.6167, 269m), 5 xi 1980, leg. David Robinson & MJW, det. MJW [ FSCA]; 2 ♂♂, 4 ♀♀, same data but clearing, forest edge, swamp & lakeshore, 24–26 vii 1977, leg. & det. DRP [ DRP].

Micrathyria mengeri Ris, 1919 . Publ. Recs. ECU: * Dunkle (1995), (H06), (P11), (H14). Also BRA, CEN, COL, GUF, GUY, PER, SUR, TTO, VEN: (P11).

Addl. Data. SU, Shushufindi   GoogleMaps : 2 ♂♂, Limoncocha, swamp, (-0.4000, -76.6167, 269m), 26 vii 1977, leg. & det. DRP [ DRP]; 3 ♂♂, same data but swamp channel, 25 viii 1980, leg. & det. SWD [ RWG, SWD].

Micrathyria occipita Westfall, 1992 . Publ. Recs. ECU: (H06), (P11), (H14); NA: Tennessen (1996b); OR: * Westfall (1992, Holotype & Allotype, 1 cop. pair, ”Lake Taracoa near Primavera” = [Francisco de Orellana], [La Nueva Esperanza] [across Río Napo from] near Primavera “Limoncocha” Río Napo & Lake Taracoa (-0.45, -76.77, 300m), 26 viii 1980, leg. SWD [FSCA], Bridges (1993), von Ellenrieder & Garrison (2011). Also PER: (H06), (P11); plus VEN: (P11).

Addl. Data. NA, Tena: [FSCA, UTIC, KJT, QZAC, TWD, USNM]; OR: Francisco de Orellana [DRP, FSCA, JJD, KJT, KWN, LSAM, QZAC, RWG, TWD, USNM]; SU, Lago Agrío: [DRP, FSCA, KJT, USNM]; Shushufindi: [FSCA, KJT, QZAC, RWG, USNM]; ZC, Zamora: [JJD].

Note: An east slope species; specific data for most records can be found in Westfall (1992).

Micrathyria ocellata Martin, 1897 . Publ. Recs. ECU: Muttkowski (1910), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997b—as M. ocellata ocellata ), (H06), (P11), (H14); CR: Calvert (1906), GY: * Martin (1897 — TYPE, Guayaquil, date?); Bridges (1993), Calvert (1906), (Ris, 1911a—as M. ocellata ocellata ), EO: Campos (1927); NA: Tennessen (1996b). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PER, SUR, TTO, VEN: (H06), (P11).

Addl. Data. EO: Machala: [KJT]; Quininde: [KWK, FSCA]; Santa Rosa [JTJ, KJT]; GY, Daule: [USNM]; Guayaquil: [FSCA, USNM]; LR: Quevedo: [FSCA]; MN, El Carmen: [JTJ]; Pedernales: [JTJ]; MS, Santiago: [SWD]; NA, Tena: [FSCA, JJD, KJT, LSAM, QZAC, SWD, USNM]; OR, Francisco de Orellana: [KJT, QZAC]; SU. Lago Agrío: [USNM]; Shushufindi: [DRP, FSCA, KJT, KWN, SWD].

Micrathyria pseudeximia Westfall, 1992 . Publ. Recs. ECU: (H06), (P11), (H14); SU: * Westfall (1992 —description). Also BOL, BRA, CEN, COL, GUF, PRY, PER, SUR, VEN, TTO: (H06), (P11); plus ARG: (P11).

Addl. Data. LR, Buena Fé: 7 ♂♂, 1 ♀, 1 cop. pair, Santo Domingo, 47 km S, forest slough at Río Palenque Field Station (200m), 18 vii 1977, leg. & det. DRP [DRP]; 8 ♂♂, 1 ♀, paratypes, same data but 8–9 x 1988, leg. & det. SWD [SWD]; MN, El Carmen: 2 ♂♂, borrow pits about 18 km NW of El Carmen (-0.1350, -79.5797, 185m), 2 iii 2010, leg. & det. JTJ [JTJ]; NA, Tena: 1 ♂, Puerto Napo 5.3 km E on Jatun Sacha Road Río Sinde swamp and pond approx. ½ km upstream (-1.0517, -77.7544, 450m), 12 xi 1997, leg. & det. WFM [FSCA]; OR, Francisco de Orellana: 1 ♂, [Pompeya Sur] S on [Pompeya Sur-Iro Road] at km marker 31, borrow pit near Río Sabaleto (-0.60, -76.47, 238m), 17 vii 1996, leg. & det. SWD [ SWD]; 1 ♂, pond, road to Waorani Reserve   GoogleMaps , 6.2 km S of turn to Yasuni Sci   GoogleMaps . Res. Sta., 10 vi 1995, leg. & det. WFM [ FSCA]; 1 ♂, swamp, P.N.Y. (-0.6508, - 76.3558, 200m), 16 xi 1997, leg. & det. KJT [ KJT]; 1 ♂, 1 ♀, same data but leg. & det. TWD [ SWD]; 1 ♂, stream 2.3 km E of Yasuni Sci. Res. Sta., 9 vi 1995, leg. & det. SWD [ SWD]; 1 ♂, Puerto   GoogleMaps Francisco de Orellana 1.7 km W on Loreto Road from Coca-Sachas Road, stream, pools & flowing shallow marsh (-0.4367, -77.0017, 258m), 13 vi 1995, leg. & det. SWD [ SWD]; 1 ♂, pond 8.2 km W of Río Payamino, Coca   GoogleMaps , 18 xi 1997, leg. & det. JJD [ JJD]; 1 paratype ♂, [La Nueva Esperanza] [across Río Napo from], near Primavera “Limoncocha”, Río   GoogleMaps Napo & Lake Taracoa   GoogleMaps (-0.45, -76.77, 300m), 26 viii 1980, leg. & det. SWD [ SWD]; Loreto: 1 ♂, S of Loreto, forest pond, (-0.8141, -77.3371, 350m), 2 xi 2013, leg. & det. CDK [ FSCA]; PC, Quito: 2 ♂♂, Pacto Parriquia, San Jose de Mashpi, SE on Mashpi road, small marshy pond & adjacent Tilapia ponds (0.1676, -78.9024, 542m), 3 ix 2018, leg. & det. KJT [ KJT, USFQ]; SD: 1 paratype ♂, Santo Domingo de Los Colorados   GoogleMaps , 12 km E on Hwy. 30 near Hotel Tinalandia, pond with water lilies (-0.30, -79.67, 610m), 6 x 1988, leg. & det. SWD [ SWD]; 1 ♂, Al- luriquin, 7.3 km W, Hotel Tinalandia, 11 iii 1990, leg. C. Anderson, det. DRP [ DRP]; SU, Lago Agrio : 1 paratype ♂, brushy ponds, 24 km W of Lago Agrio, 20 viii 1980, leg. & det. SWD [ SWD]; 2 ♂♂, swamp, 17 km S of Lago Agrio, 19 viii 1980, leg. & det. SWD [ SWD]; Shushufindi: 2 ♂♂, Limoncocha, swamp, 26 vii 1977, leg. DRP & S. Hills, det. DRP [ DRP]; 1 paratype ♂, same data but 25 viii 1980, leg. & det. SWD [ SWD]; 1 paratype ♂, [Limoncocha], [about 40 km E], or 100 km E of Coca [on Río Napo], Lago   GoogleMaps Garzacocha, La Selva Jungle Lodge (-0.4819, -76.3733, 200m), 12 x 1988, leg. & det. SWD [ SWD].

Micrathyria romani Sjöstedt, 1918 . Publ. Recs. ECU: (H06), (P11), (H14); SU: * Dunkle (1981), Westfall (1992). Also BRA, PER: (H06), (P11).

Addl. Data. SU, Shushufindi: 3 ♂♂, Limoncocha, Río   GoogleMaps Napo (-0.4000, -76.6167, 269m), 3–16 vi 1977, leg. P. Spangler & D. Givens, det.? [ USNM]; 23 ♂♂, 2 ♀♀, same data but 1–25 xi 1980, leg. & det. MJW [ FSCA]; 1 ♂, same data but forest edge, 26 vii 1977, leg. & det. DRP [ DRP]; 43 ♂♂, same data but lake edge, 26 xii 1971, D. Pearson, det. DRP [ DRP]; 16 ♂♂, same data but 26 vii 1977, leg. DRP & S. Hills, det. DRP [ DRP]; 1 ♂, same data but 11 vii 1977, leg. & det. KWK [ SWD]; 1 ♀, same data but 29 viii 1978, det. JJD [ FSCA]; 2 ♂♂, same data but 23 viii 1980, leg. & det. SWD [ KJT]; 5 ♂♂, [Limoncocha], [about 40 km E], or 100 km E of Coca [on Río Napo], Lago Garzacocha, La Selva Jungle Lodge   GoogleMaps (-0.4819, -76.3733, 200m), 11 x 1988, leg. & det. SWD [ FSCA, SWD]; 1 ♂, same data but in forest, 6 v 1990, leg. & det. E. Smith [ SWD].

Micrathyria spinifera Calvert, 1906 . Publ Recs. ECU: * Paulson(1983), (H06), (P11), (H14). Also BOL, BRA, GUF, GUY, PER, SUR, TTO, VEN: (H06), (P11).

Addl. Data. NA, Archidona: 1 ♂, along Archidona-Loreto Road, stream, pasture to forested, (-0.7164, - 77.6640, 1185m), 25 x 2013, leg. & det. CDK [FSCA]. OR, Francisco de Orellana: 1 ♂, [La Nueva Esperanza] [across Río Napo from], near Primavera “Limoncocha”, Río Napo & Lake Taracoa (-0.45, -76.77, 300m), 27 vii 1977, leg. DRP & S. Hills, det. DRP [ DRP]; 1 ♂, same data but 4 xi 1980, leg. & det. MJW [ FSCA]; La Joya de los Sachas: 1 ♂, road at Río Napo ferry, S of Limoncocha, 17 vii 1996, leg. & det. KJT [ KJT]; SU, Shushufindi: 1 ♂, Limoncocha, Río   GoogleMaps Napo (-0.4000, -76.6167, 236m), 6 ix 1972, leg. T. Emmel, det. JTJ [ FSCA]; 1 ♂, same data but forest trail, 10 viii 1971, leg. D. Pearson, det. DRP [ DRP]; 1 ♂, same data but 24 vii 1977, leg. & det. DRP [ DRP]; 7 ♂♂, 1 ♀, same data but forest edge, 25–26 vii 1977 [ DRP]; 1 ♂, same data but lake edge, 2 x 1971, leg. D. Pearson, det. DRP [ DRP]; 1 ♂, same data but 29 viii 1978, leg. KWK, det. JJD [ KWK]; 2 ♂♂, Limoncocha, small clearing near Playaco River , 24 viii 1980, leg. & det. SWD [ SWD].

Micrathyria sympriona Tennessen, 2000 . Publ. Recs. ECU: (H06), (P04), (P11), (H14); ZC: * Tennessen (2000 — Holotype & Allotype: cop. pair: [Zamora], cutgrass marsh 5.5 km SE of Zamora (-4.17, -78.94, 914m), 5 xi 1997, leg. WFM [FSCA]). Also PER: (H06), (P11); plus COL: Palacino et al. (2017).

Addl. Data. MS, Gualaquiza: 2 ♂♂, 1 ♀, Gualaquiza (-3.40, -78.58, 800m), iv–v 1935, leg. F. Campos, det. WFM [ANSP, FSCA]; ZC, Zamora: 2 paratype ♂♂, Río Jamboe slough 4.2 km SE of Zamora, 5 xi 1997, leg.

JJD, det. KJT [ JJD]; 1 paratype ♂, 1 paratype ♀, cutgrass marsh 5.5 km SE of Zamora (914m), 5 xi 1997, leg. JJD, det. KJT [ JJD]; 6 paratype ♂♂, same data but leg. & det. KJT [ KJT]; 7 ♂♂, same data but leg. WFM [ FSCA, LSAM, QZAC, USNM]. 1 ♂, small stream 5 km N of Guaguayme Alto   GoogleMaps (-3.8778, -78.8928, 880m), 4 iv 2008, leg. & det. JTJ [JTJ].

Note: The Gualaquiza specimens were labeled M. didyma .

Micrathyria tibialis Kirby, 1897 . Publ. Recs. ECU: * Paulson (1983), (H06), (P11), (H14). Also ARG, BOL, BRA, CEN, GUY, PRY, PER, SUR, VEN: (H06), (P11); plus COL: (P&P); and GUF: (P16).

Addl. Data. OR, Francisco de Orellana: 14 ♂♂, 2 ♀♀, Anangu [ SE across Río Napo from Providencia], Lago   GoogleMaps Anangu “on Río Napo ” (-0.5213, -76.4361, 300m), 9 v 1975, leg. B. Drummond III, det.? [FSCA]; SU, Shushufindi: 1 ♀, Limoncocha, Río   GoogleMaps Napo (-0.4000, -76.6167, 269m), 8 vi 1977, leg. S. Steiner & V. Warren Jr, det.? [ USNM]; 13 ♂♂, same data but 3–16 vi 1977, leg. P. Spangler & D. Givens, det.? [ USNM]; 1 ♂, same data but 25 viii 1980, leg. & det. SWD; 1 ♂, same data but 25 viii 1980 [ SWD]; same data but 6–25 xi 1980, leg. & det. MJW [ FSCA]; 1 ♂, same data but lake edge, 22 i 1972, leg. D. Pearson, det. DRP [ DRP]; 2 ♂♂, same data but 16 ii 1972 [ DRP]; 10 ♂♂, same data but 24–26 vii 1977, leg. & det. DRP [ DRP]; 1 ♂, same data but 3 ix 1980, leg. & det. KWK [ SWD]; 1 ♂, same data but 25 viii 1980, leg. KWK, det. JJD [FSCSA].

Micrathyria venezuelae De Marmels, 1989 . Publ. Recs. ECU: *(H06), (P11). Also VEN: (H06), (P11); plus (ARG): (P16); and COL: Palacino et al. (2017).

Addl. Data. EO, Portovelo: 1 ♂, Portovelo (-3.72, -79.62, 600–650m), 5 viii 1941, leg. D. Laddey, det. JJD [FSCA]; ES, Quininde: 1 ♂, Rosa Zarate, Río Cupa (0.42, -79.42, 90m), 10 i 1972, leg. D. Poveda, det. JJD [FSCA]; GY, Guayaquil: 1 ♂, Guayaquil (-2.20, -79.90, 8m), date?, leg. N. Banks, det. JJD [FSCA]; LJ, Loja: 1 ♂, seep trib. of Río Chamba, Vilcabamba (-4.2636, -79.2172, 1555m), 31 iii 2008, leg. & det. JTJ [JTJ]; LR, Buena Fé: 1 ♂, Santo Domingo, 47 km S, forest slough at Río Palenque Field Station (200m), 18 vii 1977, leg. DRP, det. WFM [ FSCA]; 6 ♂♂, 8–9 xi 1988, leg. & det. SWD [ FSCA]; Urdaneta: 1 ♂, ditch/wooded area, 2.8 km E of Hwy. E25 from km marker 98 (-1.5217, -79.5000, 35m), 21 ii 2010, leg. & det. KJT [ FSCA]; 2 ♀♀, 5 km E of Hwy. E25 (from Formisa), small stream/secondary forest, (-0.7253, -79.4444), 23 ii 2010, leg. & det. KJT ( FSCA); MN, Manta   GoogleMaps : 1 ♂, Santa Rosa, about 60 km SW of Manta near Río Seco   GoogleMaps (-1.12, -80.88, 40m), 11 v 1946, leg. L. Alonzo, det. JJD [ FSCA]; PA, Mera   GoogleMaps : 1 ♂, small streams/trails, Los Copales, between Mera and Shell (-1.4919, -78.0725, 1070m), 20 ix 2005, leg. & det. KJT [ KJT]; 1 ♂, same data but 22 ix 2005, leg. & det. JJD [ JJD]; 1 ♂, Río Anzu   GoogleMaps , 12.0 km N of Mera (-1.4181, -78.0528, 1190m), 21 ix 2005, leg. & det. KJT [ KJT]; SD: 3 ♂♂, seeps at Hotel Colorados   GoogleMaps , Hwy. 30, about 12 km E of Santo Domingo, 30 i 1997, leg. & det. JJD [ JJD]; 2 ♂♂, rocky pool, 8 km W of Alluriquin, Hwy. 30, 31 i 1997, leg. & det. JJD [ JJD].

Micrathyria unresolved:

a. Micrathyria cf. almeidai Santos, 1945 . Publ. Recs. ECU: *(H14). Also BRA: (H06), (P16). SU, Shushufindi: 1♂, Limoncocha, no addl. data provided, data provided by WAH& DLW). According to W. A. Haber (pers. comm. 2016), “It likely is not almeidai , although it keys to almeidai in Heckman (2006) , but the appendages are different.”

b. Micrathyria cf. eximia: Kirby, 1897 . Publ. Recs. ECU: (H14). Also COL, ARG, BRA: (H06), (P16). NA, Tena: 1 ♂, Puerto Napo, 5.3 km E, Río Sinde (-1.0500, -77.7481, 450m), 16 x 1988, leg. & det. SWD [RWG]. According to Rosser Garrison (pers. comm. 2011), “Probably a new species.”

c. Micrathyria cf. surinamensis: Geijskes, 1963 . Publ Recs: GUF, SUR: (H06, P16) SU: Shushufindi: 1 ♂, La Selva, 100 km E Coca, Garzacocha   GoogleMaps (-0.4983, -76.3733, 200m), 12 x 1988, leg. & det. SWD [ RWG], data provided by RWG.

d. Micrathyria sp.: ZC, Zamora: 1 ♂, shaded roadside ditch/pool, about 10–11 km N of Guaguayme Alto   GoogleMaps (-3.8356, -78.8939, 875m), 4 iv 2008, leg. & det. JTJ [ JTJ].

e. Micrathyria spp. (H14): “Three different species all in the laevigata group which needs to be revised; one of which is a new species according to Paulson and Garrison” (Haber & Wagmer pers. comm. 2016).

FSCA

Florida State Collection of Arthropods, The Museum of Entomology

SU

Stanford University

USNM

Smithsonian Institution, National Museum of Natural History

LSAM

Louisiana State Arthropod Museum

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Odonata

Family

Libellulidae