Calleida excelsa Bates, 1892

Casale, Achille & Shi, Hongliang, 2018, Revision of the Oriental species of Calleida Latreille (sensu lato). Part 1: Introduction, groups of species, and species of six species groups (Coleoptera: Carabidae: Lebiini), Zootaxa 4442 (1), pp. -1--1: 18-20

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Calleida excelsa Bates, 1892


[4] Calleida excelsa Bates, 1892  , status resurrected

Calleida excelsa Bates, 1892: 422  (type locality: Carin Chebà [Kayin State]; lectotype deposited in MSNGAbout MSNG); Andrewes, 1921: 150. (as synonym under C. lativittis  ); Jedlička, 1963: 435 (as synonym under C. lativittis  ).

Type material examined: Lectotype, designated herein, female, “ Carin Chebà [ Burma] 900-1100 m L. Fea V XII- 88 ”, “Typus”, “ Calleida excelsa Bates  ” (handwritten by Bates), “ excelsa Bates  ”, “ Calleida excelsa Bates  (es. tip.) Bates”, “ Museo Civico di Genova ” (MSNG, fig. 36 View Figure )  . Paralectotype, one female, " Birmania, Senmigion, Fea. 28

Feb. 86", " Calleida excelsa Bates  " (handwritten by Bates); "Ex.Musaeo, H.W. Bates, 1892 " ( MNHNAbout MNHN, fig. 37 View Figure ); one female, " Naga hills 3000", "Ex.Musaeo, H.W. Bates, 1892 " ( MNHNAbout MNHN).

Non-type material examined: India: one female “ India ”, “Bowring 63.47” “ Calleida lativittis Chd.  = excelsa Bates  compared with types H.E.A.[Andrewes]” ( NHMLAbout NHML); Myanmar: one female “ Rangoon E.M. Janson” ( NHMLAbout NHML); one female “ Birmania ” ( CCAAbout CCA); Thailand: one female, “ Thailand bor. Chiang Dao env. 21.5- 4.6.1995 lgt. M. Snizek” ( CCAAbout CCA); one female, “ 2.I.1998 Thaild. Chiang Mai. Samoeng, N 018°51'45.2'' E 098°43'18.7'' ” ( NHMBAbout NHMB); one female, same data ( CCAAbout CCA); China: two teneral males, " Yunnan, Yingjiang county, Nabang township, N24.75323, E97.56032, 252m, 2008. V.31, light trap, Shi Hongliang leg." ( IZASAbout IZAS); one female " Yunnan, Yingjiang county, Tongbiguan, Laoxiangping, 450m, 2003. X.29, Qin Ruihao leg." ( SWFU); one female, " Yunnan, Ruili, Huyu, 900m, 2003. XI.5, Qin Ruihao leg." ( SWFU); one female " Yunnan, Menglun town, Xishuangbanna Tropical Botany Garden, 2007. IV.20, Cao Liangming leg." ( CAUAbout CAU).

Taxonomic note. Andrewes (1921, 1930) wrongly proposed this taxon as junior synonym of C. lativittis  , and this synonymy was accepted by Jedlička (1963) and cited by Lorenz (2005). Based on the examination of type and further material of both species, we demonstrated that C. excelsa  is a valid, very distinct species.

Diagnosis. With the character states of Calleida  of the lativittis  species group, but from other known species distinguishable by the following character combination: dorsal side with dense pubescence all through head, pronotum and elytra. Elytra metallic green, disc with reddish patch on the inner three to five intervals; all antennomeres yellowish, or 3–11 somewhat darkened; femora yellowish, blackish only at extreme apex or apical third; tibiae and tarsomeres yellowish. Head wide and depressed. Pronotum transverse-cordate, widened in the middle, with lateral furrows wide and markedly flattened.

Closely related and similar to C. lativittis Chaudoir  , but easily recognized by the head, pronotum, elytral intervals and underside distinctly pubescent, and by the elytral disc with reddish patch usually wider and blackish area of femurs much shorter.

Description. The original description provided by Bates (1892) in Latin is here completed with several additional characters.

General features as in figures 36–38 View Figure : Medium sized: L: 9.0– 10.8 mm; TL: 8.4–9.8 mm. Body elongate and slender, depressed; elytra slightly widened at the posterior fifth. Head, pronotum, elytral intervals, meso- and metasternum densely punctate-pubescent.

Colour: head, pronotum, lateral margins of elytra, epipleura, underside, antennae and legs rufous; femora blackish at apex, or blackish at apical third in some individuals; elytra metallic green or bluish green; disc with a wide reddish patch, extended from the base to apex on the inner three to five intervals.

Lustre and microsculpture: shiny; head with vanished microsculpture; pronotum mostly without distinct microsculpture, except some fine transverse meshes on disc and on middle part of the basal area; elytra with very shallow isodiametric microsculpture.

Head wide and flat, narrower than pronotum, with some oblique frontal wrinkles, smooth except several fine, numerous setiferous punctures; frontal foveae very shallow. Genae rather short, tumid; eyes large and prominent; basal three antennomeres with dense additional setae.

Pronotum transverse-cordate (PW/PL: 1.12–1.25), widened in the middle, with lateral furrows wide, markedly flattened. Disc with shallow transverse wrinkles and numerous fine setiferous punctures. Lateral margin slightly reflexed, rounded in the anterior half, only sinuate in the basal sixth, then parallel sided to the posterior angles, which are rectangular. Anterior angles effaced, each with six to eight setae. Basal foveae deep and elongate.

Elytra elongate and narrow (EL/EW: 1.76–2.02), slightly widened at the posterior fifth. Striae very deep, punctate; intervals convex, with evident setiferous punctures; eighth interval not tumid at apex. Umbilicate series of 15–17 pores along the eighth stria. Apical truncation straight; outer apical angle not thickened, rounded.

Ventral side: lateral areas of meso- and metasternum and abdominal sterna with long, evident pubescence; abdominal sternum VII with two (one on each side) setae in males, four (two on each side) setae in females; apical margin of abdominal sternum VII notched in males, straight in females.

Male genitalia: only two immature males examined, with the following characters: apical orifice opened to the left; apical lamina thickened, as long as wide, rounded at apex ( fig. 41 View Figure ).

Female genitalia (reproductive tract fig. 39 View Figure and ovipositor fig. 40 View Figure ): spermatheca digitiform, a little longer than the pedicel, with very short basal projection; surface faintly and finely whorled; spermathecal pedicel with base weakly expanded; spermathecal gland duct inserted at base of the basal projection, long and slender, about two times as long as spermatheca; glandular area strongly expanded, about one fifth as long as gland duct. Gonocoxite II of ovipositor sub-rectangular, as long as 2.5 times the basal width; base much wider than apex; inner margin with several setae, extending from the middle to apex; outer margin distinctly curved at middle, only slightly setose in the apical third; apex obliquely truncate, with irregular membranous extension on the outer part.

Geographical Distribution and habitat. Distribution range as shown in map 3 View Figure : India (Nagaland); Myanmar (Carin Chebà, Senmigion, Yangon); N Thailand (Chiang Dao, Chiang Mai); SW China: Yunnan (Yingjiang, Ruili, Xishuangbanna). In deciduous and sub-tropical forests.


Museo Civico di Storia Naturale di Genova 'Giacomo Doria'


Museum National d'Histoire Naturelle


Natural History Museum, Tripoli


Culture Collection of Algae at the University of Marburg


Naturhistorisches Museum, Basel


Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine


China Agricultural University














Calleida excelsa Bates, 1892

Casale, Achille & Shi, Hongliang 2018


Calleida excelsa

Bates 1892: 422


C. lativittis

Chaudoir Lectotypus 1872


C. lativittis

Chaudoir Lectotypus 1872