Cochylis Treitschke, 1829

Sun, Yinghui & Li, Houhun, 2013, Cochylis Treitschke in China: one new species and five new records (Lepidoptera, Tortricidae, Cochylini), ZooKeys 258, pp. 85-96 : 86-87

Cochylis Treitschke, 1829

Cochylis Treitschke, 1829: 233. Type species: Tortrix roseana Haworth, 1811.

Chochylis Duponchel, 1836: 409. [misspelling of Cochylis ]

Conchylis Sodoffsky, 1837: 93. [unjustified emendation of Cochylis ]

Pontoturania Obraztsov, 1943: 97. Type species: Cochylis defessana Mann, 1861.

Acornutia Obraztsov, 1944: 68. Type species: Tortrix nana Haworth, 1811.

Cochylichroa Obraztsov & Swatschek, 1958: 233. Type species: Eupoecilia atricapitana Stephens, 1852.

Brevicornutia Razowski, 1960: 317. Type species: Cochylis pallidana Zeller, 1847.

Longicornutia Razowski, 1960: 314. Type species: Tortrix (Cochylis) phaleratana Herrich-Schäffer, 1851 sensu Razowski, 1960 [= Cochylis epilinana Duponchel, 1842]

Neocochylis Razowski, 1960: 316. Type species: Conchylis calavrytana Rebel, 1906.

Paracochylis Razowski, 1960: 316. Type species: Cochylis amoenana Kennel, 1899.

Diagnostic characters.

Cochylis is characterized by the combination of the following characters: adult small to medium; forewing with all veins separate, Sc reach ing middle of the costal margin, basal distance between R1-R2 about three times of between R2-R3, R5 to the costal margin; hindwing with costal fold in male, Rs and M1 arising from the same point or long stalked, M3 and CuA1 separate, female hindwing usually with two spines in the frenulum; male genitalia: tegumen short and broad; uncus and gnathos absent; socius separate, drooped, connected with distal part of tegumen at base; median process of the transtilla mostly developed and dentate distally (absent in a few species); sacculus developed with a hook-shaped process basally in some species, with or without terminal process; slender vinculum separate ventrally; phallus without cornutus or with cornutus composed of a bundle of spines; female genitalia: sterigma varied in size and sclerotized diversely; antrum developed and heavily sclerotized; ductus bursae short, indistinctly distinguished from the corpus bursae; corpus bursae membranous, densely suffused with tiny spines.

Biology.

In the Palaearctic Region, there are one or two generations annually, and over-wintering occurs in the larval stage. Larvae are oligophagous and feed mainly on plants belonging to Asteraceae ( Razowski 1987).

Distribution.

All species are distributed in the Holarctic, Oriental and Neotropical regions.

Key to Chinese species of Cochylis based on male genitalia

1	Phallus uniformly slender	2
-	Phallus stout basally, slender distally	6
2	Phallus with cornutus ( Razowski 1970: Taf. 106, Fig. 270)	Cochylis atricapitana
-	Phallus without cornutus	3
3	Transtilla without median process ( Fig. 7 )	Cochylis discerta
-	Transtilla with median process	4
4	Phallus slightly sinuate ( Razowski 1970: Taf. 104, Fig. 260)	Cochylis roseana
-	Phallus curved orthogonally distally	5
5	Sacculus extended to a hook-shaped process basally ( Razowski 1970: Taf. 105, Fig. 2681-4)	Cochylis hybridella
-	Sacculus not extended basally ( Fig. 8 )	Cochylis dubitana
6	Transtilla without median process ( Razowski 1970: Taf. 109, Fig. 283)	Cochylis psychrasema
-	Transtilla with median process	7
7	Valva with a spine-shaped process at basal 1/3 near outer margin ( Fig. 11 )	Cochylis triangula sp. n.
-	Valva without process near out margin	8
8	Median process nearly triangular ( Fig. 10 )	Cochylis posterana hyrcana
-	Median process short stripe-shaped, nearly parallel-sided	9
9	Socius about 1/2 length of median process ( Razowski 1970: Taf. 108, Fig. 279)	Cochylis defessana
-	Socius almost same length as median process	11
11	Valva with outer margin slightly protruded distally, costa straight ( Fig. 9 )	Cochylis faustana
-	Valva with outer margin nearly straight distally, costa slightly concave ( Razowski 1970: Taf. 108, Fig. 2771-2)	Cochylis piana

Figures

Figures 7 - 10. Male genitalia of Cochylis spp. 7 Cochylis discerta Razowski, slide No. SYH 11644 8 Cochylis dubitana (Huebner), slide No. SYH 10244 9 Cochylis faustana (Kennel), slide No. SYH 11447 10 Cochylis posterana hyrcana (Toll), slide No. SYH 11434 11 Cochylis triangula sp. n., paratype, slide No. SYH 10200. (scales: 0.5 mm).