Caryophyllia arcotensis ( Forbes, 1846 )

Baron-Szabo, Rosemarie C., 2008, Dendrophylliina, Caryophylliina, Fungiina, Microsolenina, and Stylinina, Zootaxa 1952, pp. 1-244 : 45-48

Caryophyllia arcotensis ( Forbes, 1846)

Pl. 4, figs 1a, b, Text-Fig. 1

v*1846 Turbinolia arcotensis , sp. nov.: Forbes, p. 163, pl. 19, figs 9a–b.

1850 Ellipsosmilia arcotensis: d’Orbigny , vol. 2, p. 276.

1851b Trochosmilia arcotensis: Milne Edwards & Haime , p. 17.

non1873 Caryophyllia arcotensis, Stoliczka : Stoliczka, p. 7, pl. 1, figs 1–10.

1873 Turbinolia arcotensis, Stoliczka , p. 8, pl. 1, figs 11–12.

non1922 Caryophyllia arcotensis, Forbes : Cottreau, p. 169, pl. 1, figs 6–6b.

non1936 Caryophyllia arcotensis Forbes : Alloiteau, p. 9.

(v)1951b Caryophyllia arcotensis Stol. , var. regularis nov. var.: Alloiteau, p. 81, pl. 13, figs 1–1a.

(v)1951b Caryophyllia antonibensis , nov. sp.: Alloiteau, p. 80, pl. 13, figs 2–4.

1955 Caryophyllia antonibensis All. : Rossi Ronchetti, p. 113, pl. 11, figs 5–6.

non1958 Protrochocyathus arcotensis Forbes , sp.: Alloiteau, p. 136, pl. 17, figs 6a–g.

non1995 Caryophyllia arcotensis ( Forbes, 1846) : Tchéchmédjiéva, p. 55, pl. 10, figs 11, 12.

2000 Caryophyllia antonibensis, Alloiteau 1951 : Löser, p. 17.

2000 Caryophyllia arcotensis ( Forbes 1846) : Löser, p. 17.

PLATE 4:

Fig. 1 Caryophyllia arcotensis ( Forbes, 1846) , holotype, BMNH, R.29211, Maastrichtian of India, scale bar: 4.5 mm. 1a, cross view; 1b, longitudinal view. Fig. 2 Caryophyllia mediavia Vaughan & Popenoe, 1935 , holotype, NMNH, M371041, Danian of the USA, scale bar: 3.4 mm. 2a, cross view; 2b, longitudinal view. Fig. 3 Bathycyathus lloydi ( Vaughan, 1920) (holotype of Steriphonotrochus manorensis, Wells, 1933 ), NMNH, Wells coll., I75132, Upper Maastrichtian of the USA. 3a, upper surface, cross view, scale bar: 5 mm; 3b, polished surface, juvenile stage of corallum, scale bar: 1 mm; 3c, longitudinal view, scale bar: 2 mm. Fig. 4 Bathycyathus lloydi ( Vaughan, 1920) , holotype, NMNH, Vaughan coll., I32376, Paleocene of the USA, scale bar: 2.5 mm. 4a, upper surface, cross view; 4b, longitudinal view. Fig. 5 Trochocyathus speciosus ( Gabb & Horn, 1860) (syntype of Trochocyathus epicharis Wanner, 1902 ), NRM-PZ, Wanner coll., Cn 47525, Danian of Egypt, scale bar: 1.5 mm. 5a, cross view, upper surface; 5b, longitudinal view; 5c, side view.

Dimensions. d (min)=16.5– 24 mm; d (max)= 17–25 mm; d (min)/d (max)=0.8–1; s=40–48+s5; h=up to 35 mm.

Description. Solitary, variable conical (e.g., turbinate, cylindrical), slightly elliptical or circular in outline; costosepta straight, developed in 4 complete cycles in 6 systems, regularly or irregularly alternating in length and thickness; septa laterally smooth or finely granulated; pali seem to be axial to the second set of septa.

Text-Fig. 1 Caryophyllia arcotensis ( Forbes, 1846) as figured in Alloiteau (1951b) (as holotype of Caryophyllia arcotensis var. regularis Alloiteau, 1951b ). A, B. Upper Maastrichtian of Madagascar, scale bar: 10 mm; A, upper surface longitudinal view; B, cross view of corallum.

Remarks. The specimens described from the Albian of Madagascar as Caryophyllia arcotensis ( Forbes, 1846) by Alloiteau (1936), which were later transferred to the genus Protrochocyathus in Alloiteau, 1958 ( Protrochocyathus is interpreted here to be the junior synonym of Trochocyathus ), have calicinal diameters ranging from 8–10 mm with 48 septa arranged in 6 systems. Therefore, they most likely represent junior syn-

The specimens described as Caryophyllia arcotensis (Forbes) from the Upper Campanian of Bulgaria by Tchéchmédjiéva (1995) differ from the genus Caryophyllia in lacking pali. They most likely belong to the genus Parasmilia .

Type locality of species. Maastrichtian of India (Pondicherry) .

Distribution. Maastrichtian of India and Libya (Sofeggin), Upper Maastrichtian of Madagascar.