Palaeopsammia zitteli ( Vaughan, 1900 )

Baron-Szabo, Rosemarie C., 2008, Dendrophylliina, Caryophylliina, Fungiina, Microsolenina, and Stylinina, Zootaxa 1952, pp. 1-244 : 38-39

Palaeopsammia zitteli ( Vaughan, 1900) non Wanner, 1902

Pl. 1, figs 4a, b, Pl. 2, figs 1–3b, 5a–8b, Pl. 3, figs 2a, b

v1897 Placosmilia n. sp.: Merriam, p. 770.

v*1900 Trochocyathus zitteli Merriam : Vaughan, p. 102, pl. 8, figs 4–6a, fig. 7.

v1902 Thecocyathus aegyptiacus n. sp.: Wanner, p. 99, pl. 14, figs 1–1a.

v1902 Palaeopsammia zitteli n. sp.: Wanner, p. 104, pl. 15, figs 1–4.

1902 Caryophyllia Jasmundi n. sp.: Wanner, p. 100, pl. 14, figs 2–4.

v1902 Palaeopsammia multiformis n. sp.: Wanner, p. 105, pl. 15, figs 5–9.

1902 Palaeopsammia zitteli Wanner : Quaas, p. 161.

1903 Palaeopsammia zitteli Wanner : Oppenheim, p. 448, pl. 7, figs 17–18a.

v1920 Steriphonotrochus leithensis Vaughan , n. sp.: Vaughan, p. 64, pl. 10, figs 6–6b.

1925 Trochocyathus zitteli Vaughan 1900 : Felix, pars 28, p. 204.

v1931 Microseris complanata: Collignon , pl. V (I), figs 4–5b.

v1933 Palaeopsammia fastigiata nov. sp.: Kühn, p. 183, pl. 17, fig. 9.

?1955 Trochocyathus cf. epicharis Wann. : Rossi Ronchetti, p. 108.

v1958 Microseris complanata Collignon : Alloiteau, p. 153.

v1958 Diegosmilia complanata Collignon sp.: Alloiteau, p. 153, pl. 34, figs 5, 6, text-fig. 25.

v1958 Kumbiopsammia Besairiei nov. sp.: Alloiteau, p. 201, pl. 34, figs 1–4.

1958 Palaeopsammia multiformis Wanner : Alloiteau, p. 203.

1958 Palaeopsammia zitteli Wanner : Alloiteau, p. 203.

v1958 Dasmosmilia ? spissa n. sp.: Squires, p. 41, pl. 6, figs 15–16.

1970 Palaeopsammia multiformis Wanner : Hassan & Salama, p. 92, pl. 3, figs 1a–j.

1970 Kumbiopsammia cf. besairiei Alloiteau : Hassan & Salama, p. 94, pl. 3, figs 2a–i.

1971 Palaeopsammia multiformis Wanner : Salama, p. 86, pl. 10, figs 1–5.

1981 Palaeopsammia multiformis Wanner : Barthel & Herrmann-Degen, p. 162.

1987 Palaeopsammia multiformis Wanner, 1902 : Kuzmicheva, p. 146, pl. 15, figs 7–9.

(v)1995 Palaeopsammia multiformis Wanner 1902 : Abdel-Gawad & Gameil, p. 27, pl. 20,figs 5–6.

1995 Bathypsammia cleopatrae Youssef and Salama, 1969 : Abdel-Gawad & Gameil, p. 28, pl. 20,figs 9–10. non1996 Caryophyllia jasmundi Wanner, 1902 : Schuster, p. 80.

(v)1996 Notocyathus (Notocyathus) tineidensis n. sp.: Schuster, p. 81, pl. 19, figs 3a, b, text-fig. 43.

(v)1996 Palaeopsammia multiformis Wanner, 1902 : Schuster, p. 83, text-fig. 46.

(v)1996 Kumbiopsammia salamae n. sp.: Schuster, p. 84, text-fig. 48.

1996 Kumbiopsammia cf. besairiei Alloiteau, 1958 : Schuster, listed in appendix.

v1997 Palaeopsammia multiformis Wanner, 1902 : Cairns, p. 11 (older synonyms cited therein).

2000 Dasmosmilia spissa Squires : Löser, p. 27.

2000 Diegosmilia complanata ( Collignon 1931) : Löser, p. 28.

v2000 Diegosmilia complanata ( Collignon, 1931) : Baron-Szabo, p. 126, pl. 12, fig. 1.

2000 Caryophyllia jasmundi, Wanner 1902 : Löser, p. 17.

2000 Kumbiopsammia besairiei, Alloiteau 1958 : Löser, p. 44.

2000 Palaeopsammia fastigiata, Kühn 1933 : Löser, p. 58.

2000 Palaeopsammia zitteli, Wanner 1902 : Löser, p. 58.

2001 Palaeopsammia aegyptiaca (Wanner, 1902) : Cairns, p. 13.

v2001 Kumbiopsammia besairiei Alloiteau, 1958 : Cairns, p. 40, pl. 14, figs g–i.

v2002 Diegosmilia complanata ( Collignon, 1931) : Baron-Szabo, p. 16, pl. 119, figs 2–3, 6.

2002 Palaeopsammia fastigiata Kühn, 1933 : Baron-Szabo, p. 78.

2002 Palaeopsammia zitteli Wanner, 1902 : Baron-Szabo, p. 78.

v2002 Kumbiopsammia besairiei Alloiteau, 1958 : Baron-Szabo, p. 79, pl. 57, fig. 2

ken piece).

Description. Solitary, variably conical or patellate; costosepta arranged in 4–5 cycles in 6 systems in corallites of around 15 mm and larger; S1–S3 nearly equal in adult corallites, reaching the axial region; S1 often branch near the outer edge; oldest septa often straight, younger ones tend to be irregularly wavy; columella generally well-developed, spongy, fasciculate or formed by twisted segments, sometimes appearing net-like; columella projecting or deeper in corallum.

Remarks. The specimens documented as Bathypsammia cleopatrae Youssef & Salama, 1969 , in Abdel- Gawad & Gameil (1995) from the Paleocene of Egypt differ from the generic concept of Bathypsammia Marenzeller, 1906 , in having a rather patellate shaped, free corallum, and lacking the septal arrangement according to the Pourtalès plan, thus closely agree with Palaeopsammia Wanner, 1902 .

In his revision on dendrophylliids Cairns (2001, p. 13) merged the species Caryophyllia jasmundi and Trochocyathus aegyptiacus and transferred them to the genus Palaeopsammia . The material documented in Schuster (1996) however, seems to closely correspond to the genus Caryophyllia .

The specimens in the above list are interpreted to belong to the same species based on the occurrence of identical septal arrangement in corallites of the same size. The fact that in some specimens all of the septa are very wavy and flexuous whereas in others the septa seem rather straight is interpreted as a taxonomically irrelevant factor since in some specimens of the Egyptian type collection both types of septa appear, sometimes they are straight only in their uppermost margins but are flexuous a few mm below (see e.g., Pl. 2, Fig. 5a, upper part of image). In addition, in the Egyptian specimens the corallite walls are often very poorly preserved and can only be observed in a few places, which can result in appearances that seem to be different from the specimens with well-preserved wall structures. Moreover, in some corallites the columellar structures are more bulging than in others, e.g., the specimen figured on Pl. 2, Fig. 7b shows axial developments that seem to be different from the ones seen in the specimen figured on Pl. 2, Fig. 6. However, in both corals the columella is well-developed spongy-papillose. Moreover, in the Egyptian type material, in addition to specimens having a bulging columella, forms showing a flat or depressed columella occur (e.g., specimen figured on Pl. 2, Fig. 8a has a rather flat, in some places even depressed, columella). Therefore, these specimens are considered synonymous to the same species.

Type locality of species. Danian of the USA ( Martinez Group , California) .

Distribution. Cretaceous of Egypt, (?)Upper Cretaceous-(?)Eocene of New Zealand, Cenomanian and Maastrichtian of Madagascar, Maastrichtian of Mexico,? Libya, Iran (Neyriz, Esfahan), Tunisia, and Turkmenistan, Maastrichtian-Paleocene of Egypt, Middle-Upper Maastrichtian of the UAE / Oman border region, Paleocene of Libya and the USA (Cannonball Marine Formation, South Dakota; Martinez Group, California).

New Material. Danian of Egypt, NRM-PZ, sample nos.: Cn 47505–47515 and 47518–47565 (Babel Jasmund).