Ebenomitra januszkiewiczi (Friedberg, 1928) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4982 (1), pp. 1-70 : 33-34

Ebenomitra januszkiewiczi (Friedberg, 1928) nov. comb.

Figs 10G 1 –G 2 , H 1 –H 2 , I 1 –I 2 , J

Mitra sp. ign; an nova.— Friedberg 1911: 31, text-fig. 7.

Mitra Januszkiewiczi Friedb. —Friedberg 1928: 578, pl. 37, fig. 25.

Mitra Januszkiewiczi Friedb. — Friedberg 1938: 130.

Mitraria (Mitraria) hilberi ( Hoernes & Auinger, 1880) —Bałuk 2006: 215, pl. 14, fig. 8 [= non Hoernes & Auinger, 1880].

Type material. Lectotype (designated herein): ZNG PAN A-I-50/35, SL: 14.0 mm, MD: 5.1 mm, Zboriv ( Zborów ) ( Ukraine), stored in the Geological Museum of the Institute of Geological Sciences , Polish Academy of Sciences, illustrated in Friedberg (1928, pl. 37, fig. 25), figs 10G 1 –G 2 . Paralectotypes: ZNG PAN A-I-50/36.1–36.6, SL: 15.2 mm, MD: 5.1 mm, Zboriv ( Zborów ) ( Ukraine), stored in the Geological Museum of the Institute of Geological Sciences, Polish Academy of Sciences, figs 10H 1 –H 2 ; ZNG PAN A-I-50/36.1–36.6, SL: 9.1 mm, MD: 3.5 mm, Zboriv ( Zborów ) ( Ukraine), stored in the Geological Museum of the Institute of Geological Sciences, Polish Academy of Sciences, figs 10I 1 –I 2 ; ZNG PAN A-I-50/36.1–36.6, SL: 12.1 mm, MD: 4.8 mm, Zboriv ( Zborów ) ( Ukraine), stored in the Geological Museum of the Institute of Geological Sciences, Polish Academy of Sciences, fig. 10J .

Revised description. Shell moderately large, moderately slender pupiform, with high, weakly gradate spire. Protoconch paucispiral with broadly convex, low nucleus and about one weakly convex whorl. Transition to teleoconch indistinct in available specimens. Teleoconch of six glossy, weakly convex whorls, rapidly increasing in height, with periphery in abapical third, separated by deeply impressed, narrowly canaliculated suture. No axial sculpture on spire whorls. Last whorl weakly convex to subcylindrical, moderately constricted at short base, bearing subobsolete axial ribs, most prominent along periphery and base. Aperture wide, ovoid, anal canal indistinct with weak parietal denticle. Columella with three to four columellar folds. Adapical two folds prominent, abapical folds weak, fourth fold only present in one specimen. Outer lip thin, with about 12 long, delicate lirae within. Siphonal canal short, straight or slightly bent to left, with wide siphonal notch.

Shell measurements and ratios. SL: 9.1–15.2 mm, MD: 3.5–5.1 mm; AA: 25°, AA: 45–52°, SL/MD: 2.9–3.1, AL/AW: 3.0–3.4, AH/S: 3.0.

Discussion. We place this rare species in Ebenomitra based on its paucispiral protoconch, which excludes a placement in Pusia (see Fedosov et al. 2017). Nevertheless, the high spire and canaliculate suture are atypical for Ebenomitra and the occasional presence of a fourth columellar fold is also problematic. The species is somewhat reminiscent of Ebenomitra pseudopyramidella ( Boettger, 1906) , but differs in its higher spire, more slender profile, deeper suture, and lacks axial sculpture on early spire whorls. The same features allow a separation from Pusia avellanella ( Boettger, 1906) , which is similar concerning the axial ribs along the periphery, but that species is immediately separated by its multispiral protoconch, which places it in the genus Pusia Swainson, 1840 , rather than Ebenomitra .

Palaeoecology. The mollusc assemblage from the type locality suggest shallow marine near-shore environments (own data).

Distribution in Central Paratethys. Badenian (middle Miocene): Korytnica Basin: Korytnica ( Poland) (Bałuk 2006); Ukrainian Fore-Carpathian Basin: Zborów (Zboriv), ( Ukraine) (Friedberg 1928).

Figures

FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 10 A1–A2. Ebenomitra leucozona (Andrzejowski, 1830), Lăpugiu de Sus (Romania), NHMW 1870/0033/0034. B1– B2. Ebenomitra leucozona (Andrzejowski, 1830), Gainfarn (Austria), NHMW 1846/0037/0110. C1–C2. Ebenomitra leucozona (Andrzejowski, 1830), Lăpugiu de Sus (Romania), NHMW 1868/0001/0401. D1–D2. Ebenomitra pseudopyramidella (Boettger, 1906), Ternopil (Ukraine), NHMW 2020/0071/0003. E1–E2. Ebenomitra pseudopyramidella (Boettger, 1906), holotype, Ternopil (Ukraine), NHMW 2020/0071/0001. F1–F2. Ebenomitra pseudopyramidella (Boettger, 1906), Ternopil (Ukraine), NHMW 2020/0071/0002. G1–G2. Ebenomitra januszkiewiczi (Friedberg, 1928), lectotype, Zboriv (Zborów) (Ukraine), PAN A-I-50/35. H1–H2. Ebenomitra januszkiewiczi (Friedberg, 1928), paralectotype, Zboriv (Zborów) (Ukraine), ZNG PAN A-I-50/36.1-36.6. I1–I2. Ebenomitra januszkiewiczi (Friedberg, 1928), paralectotype, Zboriv (Zborów) (Ukraine), ZNG PAN A-I-50/36.1-36.6. J. Ebenomitra januszkiewiczi (Friedberg, 1928), paralectotype, Zboriv (Zborów) (Ukraine). ZNG PAN A-I-50/36.1-36.6.