Entada glandulosa Pierre ex. Gagnep., Notul. Syst. (Paris) 2: 57. 1911.

O'Donnell, Shawn A., Ringelberg, Jens J. & Lewis, Gwilym P., 2022, Re-circumscription of the mimosoid genus Entada including new combinations for all species of the phylogenetically nested Elephantorrhiza (Leguminosae, Caesalpinioideae, mimosoid clade), PhytoKeys 205, pp. 99-145 : 99

Entada glandulosa Pierre ex. Gagnep., Notul. Syst. (Paris) 2: 57. 1911.

= Entada tamarindifolia Pierre ex. Gagnep., Notul. Syst. (Paris) 2: 59. 1911.

Type.

LAOS. Massie s.n. (lectotype: P [P02436137], designated by I.C. Nielsen in Adansonia ser. 2, 19: 342. 1980) .

Description.

Shrub, scandent (Fig. 14A ). Leaves: petiole 1.8-4 cm long, rachis 4.5-10 cm long, terminating in a bifurcating tendril; pinnae 2 pairs pair leaf, 4-8 cm long, with 5-6 pairs of leaflets; leaflets 1.1-4 × 0.5-1.7 cm, elliptic to oblong, base truncate, apex emarginate or mucronate. Inflorescence: a spike 7-18 cm long, axillary, solitary, rachis pubescent to velutinous (Fig. 14A, B ). Flowers: creamy white to yellowish-white, sub-sessile; calyx cupular, 2-2.5 mm long, glabrous to puberulous; petals lanceolate, 5 × 1 mm, a pair of linear glands on the lower half of the dorsal side of each petal; stamen filaments 8 mm long (Fig. 14B ). Fruit: a torulose, curved craspedium, 35 × 2.2-2.6 cm, with transverse septa between seeds dividing the fruit into one-seeded segments which, upon ripening, fall from the persistent replum; segments 2.4 cm long; epicarp coriaceous, endocarp papyraceous (Fig. 14C ). Seeds: sub-globular, 1.1-1.8 cm, hard, brown, pleurogram lacking.

Distribution.

Laos, Cambodia, southern Vietnam, Thailand, Myanmar.

Habitat and ecology.

Seasonally dry deciduous forest, mixed forest with Dipterocarpaceae and evergreen forest, up to 500 m alt. Usually on limestone, though also in shallow sandy soils and in red soils.

Figures

Figure 14. Entada glandulosa habit and reproductive structures. A scandent shrub with erect axillary spikes, Thailand (photo: T Boonkerd, all rights reserved) B solitary axillary spikes with open flowers, Thailand (photo: T Boonkerd, all rights reserved) C immature fruits, Thailand (photo: T Boonkerd, all rights reserved).