Leseha Smith, 2019

Smith, David R. & Nishida, Kenji, 2019, A new genus and three new species of Neotropical sawflies (Hymenoptera, Tenthredinidae) from Costa Rica, with host plants and life history notes, Journal of Hymenoptera Research 72, pp. 45-65 : 45

Leseha Smith gen. nov.

Type-species.

Leseha vespa Smith & Nishida

Description.

Antenna with scape and pedicel each longer than broad. Mandibles evenly curved, each with one subapical tooth. Clypeus truncate to very shallowly emarginate. Malar space less than half diameter of an ocellus. Genal carina absent. Eyes large, converging below, lower interocular distance subequal to or slightly less than eye height. Head from above narrowing behind eyes; distance behind eyes half or less eye length. Epicnemium elongate, flat, on same level as mesepisternum and separated by a fine suture. Anal cross vein of forewing absent. Anal cell of hindwing sessile. Tarsal claws with one inner tooth about as long and broad as outer tooth and located near outer tooth; basal lobe absent.

Etymology.

The genus name is an arbitrary combination of letters; the gender is feminine.

Remarks.

Leseha is separated from other genera of Selandriinae by the bifid tarsal claw without a basal lobe ( Fig. 53 ), mandibles each with one inner tooth, absence of a genal carina, and elongate, flat epicnemium. It is similar to Adiaclema , but Adiaclema has simple mandibles. The claws and epicnemium are similar to Liliacina , but Liliacina has a deeply circularly emarginated clypeus and mandibles are long and bent at almost a right angle. In existing keys ( Smith 2006) it will key to Andeana in couplet 22; however, the following will separate Leseha from Andeana (characters in parentheses): tarsal claws ( Fig. 53 ) with long inner tooth, about as long and broad as outer tooth and situated next to outer tooth (inner tooth small, less than half length of outer tooth and situated near center of claw, as in Fig. 54 ); eyes large ( Figs 17 , 39 ), lower interocular distance subequal to or slightly less than eye height (small, lower interocular distance greater than eye height, as in Fig. 51 ); head in dorsal view ( Figs 18 , 40 ) narrowing behind eyes with distance behind eyes less than half eye length (head in dorsal view straight behind eyes, distance behind eyes more than half eye length, as in fig. 52); anal cell of hindwing sessile, as in Fig. 1 (anal cell of hindwing with short petiole, as in Fig. 55 ).

Hosts recorded for other Selandriinae are Entodontopsis leucostega (Brid.) W.R. Buck and Irel. ( Stereophyllaceae ) for Adiaclema chigiyae Smith ( Smith and Janzen 2003b); Blechnum occidentale L. ( Blechnaceae ) for Dochmioglene sp., and Lomariopsis vastita E. Fourn. ( Lomariopsidaceae ) for Dochmioglene crassa (Cameron) ( Smith et al. 2013).

Species included.

In addition to the two new species described here, the following four South American species also belong in this new genus.

Leseha cruralis (Konow), comb. nov.

Stromboceros cruralis Konow 1899: 296. ♀. "Peru (Callanga, Cuzco)"; Konow 1905: 98; Oehlke and Wudowenz 1984: 376 (holotype); Taeger et al. 2010: 532.

Material. Bolivia, Ecuador, Peru.

Notes. Two females at SDEI are labeled as types: the one labeled "Callanga, Cuzco, Peru," "Coll. Konow," “TYPE,” " Stromboceros cruralis Knw., Peru" is here designated lectotype. The other specimen from "Pachitea Peru" is not part of the type series since the locality was not mentioned by Konow.

Leseha cubitalis (Malaise), comb. nov.

Dochmioglene cubitalis Malaise 1954: 283. ♀, ♂. no locality in original description; Taeger et al. 2010: 487.

Material. Brazil (Santa Catarina).

Notes. Malaise (1954) did not designate a holotype; he stated "(10 ♀♀, numerous ♂♂)", so the exact number of specimens he had is unknown. The lectotype, here designated, is labeled "Brasilien, Nova Teutonia, 27°11'B., 52°23'L, Fritz Plaumann," "11 Mar. 1942," “TYPUS,” " Dochmioglene cubitalis sp. nov., R. Malaise det. 1952," "31 71," "Riksmuseum Stockholm" (NHRS).

Leseha joergenseni (Schrottky), comb. nov.

Stromboceros joergenseni Jörgensen 1913: 274. ♀, ♂. nomen nudum.

Siobla joergenseni Schrottky 1913: 702. ♀, “♂”? “Bompland”; Taeger et al. 2010: 540; Abrahamovich et al. 2010: 67 (two syntypes, male and female, at LP).

Material. Argentina, Brazil.

Notes. Jörgensen’s mention of this species predates Schrottky’s description, but Jörgensen gave no description, only "Junto con la especie anterior, pero acaso no tan comun." Schrottky stated “♀♂,” "22-IX-09, Bompland (Jorgensen Nr. 3)." DRS saw three females from La Plata (LP), no males. There are two females on the same pin with a label "Argentina, Bonpland, 22-IX-1909, P. Jorgensen," " Siobla joergenseni Schrottky, C. Schrottkky det. 1012." The lectotype, here designated, is the top specimen on this pin. The other female DRS saw has a different date, 20-IX-1910, and even though labeled as Siobla joergenseni , it belongs to the genus Plaumanniana .

Leseha limbata (Kirby). comb. nov.

Selandria limbata Kirby 1889: 141. ♀, ♂. "Theresopolis, Brazil"; Dalla Torre 1894: 143; Taeger et al. 2010: 540.

Stromboceros limbatus : Konow 1905: 98.

Material. Brazil (Rio de Janeiro, Santa Catarina).

Notes. The holotype, at BMNH, is a male labeled "Type H.T.," "B.M. Type Hym. 1.303," "B. M. Type Hym., Selandria limbata ( Kirby 1889)," "limbata type," “27/987,” "Theresopolis, 88-137." DRS could not find the female.

Leseha strigata (Enderlein), comb. nov.

Strongylogaster strigatus Enderlein 1920: 363. ♀. “Südbrasilien, Santa Catarina; Brasilien, Espiritu-Santo"; Taeger et al. 2010: 537.

Material. Brazil (Santa Catarina).

Notes. Enderlein saw two females, one from each of the localities, but DRS saw only one female from Santa Catarina. It is labeled "S. Catarina, Lüderwaldt,” “Type,” " Stromboceros strigatus Endl., type F, Dr. Enderlein det 1918," Muz. Zool. PAN Warszawa 12/75" and is here designated the lectotype.

Figures

Figures 51 - 55. 51, 52, 54, 55 Andeana sp. 53 Leseha vespa 51 head front 52 head dorsal 53 trsal claw 54 tarsal claw 55 hindwing.

Figures 16 - 23. Leseha vespa 16 adult female, ovipositing 17 head, front 18 head, dorsal 19 sheath, lateral 20 female lancet 21 apical tergite of male 22 male genital capsule 23 male penis valve.

Figures 38 - 44. Leseha carranzae 38 female, dorsolateral view 39 head, front 40 head, top 41 female sheath, lateral 42 female lancet 43 male genital capsule 44 male penis valve.

Figures 1 - 8. Waldheimia saurauia (except 6) 1 dorsal view, female 2 head, front 3 head, dorsal 4 female sheath, lateral 5 female lancet 6 female lancet of W. laeta 7 male genital capsule 8 male penis valve.