Exocelina yoginofi Shaverdo & Balke

Shaverdo, Helena, Sagata, Katayo & Balke, Michael, 2018, Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 803, pp. 7-70: 40-43

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Exocelina yoginofi Shaverdo & Balke

sp. n.

24. Exocelina yoginofi Shaverdo & Balke  sp. n. Figs 22, 46

Exocelina  undescribed sp. MB1302: Toussaint et al. 2014: supplementary figs 1-4, table 2; Toussaint et al. 2015: supplementary figs S1-S2, table S3.

Exocelina yoginofi  _ New_Guinea_MB1302: Toussaint et al. 2015: supplementary information S5-S6.

Type locality.

Papua New Guinea: Eastern Highlands Province, Kainantu, Yoginofi, 06°21.80'S, 145°45.46'E, 1,900 m a.s.l.

Type material.

Holotype: male "Papua New Guinea: Eastern Highlands, Kainantu, Yoginofi, 1900m, 9.v.1994, 06.21.799S 145.45.463E, Balke & Sagata (PNG 55)" ( ZSM). Paratypes: 6 males, 8 females with the same label as the holotype, one male with an additional green label "M.Balke 1302" ( NHMW, ZSM). 1 male, 1 female "Papua New Guinea: Eastern Highlands, 37 km S Goroka, Hogave vill., Mt. Michael, 2179-2800m, 9.-15.vii.2009, 06.22.479S 145.15.256E, Sagata (PNG230)" ( ZSM).


Body size and form: Beetle medium-sized: TL-H 3.45-4.0 mm, TL 3.85-4.45 mm, MW 1.85-2.15 mm (holotype: TL-H 3.9 mm, TL 4.3 mm, MW 2.0 mm), with oblong-oval habitus.

Coloration: Dark brown to piceous, with paler pronotum. Head reddish to dark brown, paler anteriorly and posteriorly. Pronotum brown to dark brown, with reddish brown lateral sides and sometimes anteriorly and narrowly posteriorly, darker on disc, sometimes to piceous. Elytra brown to piceous, sometimes with narrow reddish sutural lines. Head appendages and legs proximally yellowish red, legs distally darker, reddish brown (Fig. 22).

Surface sculpture: Submatt dorsally. Head with rather dense punctation (spaces between punctures 1-2 times size of punctures), finer and sparser anteriorly; diameter of punctures smaller than or equal to diameter of cells of microreticulation. Pronotum and elytra with sparser and finer punctation than on head. Head, pronotum and elytra with rather strongly impressed microreticulation. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and fine, sparse but distinct punctation.

Structures: Pronotum with distinct lateral bead. Its lateral sides with distinct longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 rounded.

Male: Antennomere simple (Fig. 22). Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 long and narrow, with narrow anterior band of ca. 40 and posterior row of 14 relatively long setae (Fig. 46D). Median lobe in lateral view slightly curved, with apex dully pointed, slightly bent downwards; in ventral view, broadened subdistally, with broad, rounded apex. Paramere very slightly concave on dorsal side and with dense setae on subdistal part; proximal setae inconspicuous (Fig. 46 A–C). Abdominal ventrite 6 with 6-8 lateral striae on each side.

Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.


Exocelina yoginofi  sp. n. is similar to E. simbaiensis  sp. n. in body form, size, and coloration, but differs from it in distinctly finer punctation and microreticulation and in having the apex of the median lobe not thickened. Also, see under E. okapa  sp. n.


Papua New Guinea: Eastern Highlands Province (Fig. 50).


The species is named after Yoginofi Village. The name is a noun in the nominative singular standing in apposition.