Andrena, Fabricius, 1775

Kratochwil, Anselm & I, Canary, 2020, Revision of the Andrena wollastoni group (Hymenoptera, Anthophila, Andrenidae) from the Madeira Archipelago and the Canary Islands: upgrading of three former subspecies and a description of three new subspecies, Linzer biologische Beiträge 52 (1), pp. 161-244 : 195-196

Andrena g. gomerensis WARNCKE, 1993

Syn. Andrena wollastoni gomerensis WARNCKE, 1993 p.p.

H o l o t y p e: 1♀ La Gomera, Chipude(s), 22.04.1942, collector unknown (ID-No OLML38, former holotype of A. wollastoni gomerensis).

P a r a t y p e s: 2♀♀ La Gomera, Costa de Chipude (s), 25.04.1942, collector unknown (ID-No OLML44 , OLML45 , former paratypes of A. wollastoni gomerensis) ; 7♀♀ La Gomera, Degollada de Peraza , 945 m, 12.05.1992, leg. J.A.W. Lucas (ID-No OLML6 - OLML12 ) ; 3♀♀ La Gomera, Igualero , 1290 m, 15.05.1992, leg. J.A.W. Lucas (ID-No OLML13 - OLML15 ) ; 1♂ La Gomera, Chipude (s), 22.04.1942, collector unknown (ID-No OLML42 , former paratype of A. wollastoni gomerensis) ; 3♂♂ Costa de Chipude (s), 25.04.1942, collector unknown (ID-No OLML40 , OLML41 , OLML43 , former paratypes of A. wollastoni gomerensis) ; 1♀ Montaña de la Zarza, 1000 m, 20 km WNW S. Sebastián , 03.06.1983, leg. Hohmann (MCNSC, HY-7031) .

Not chosen as paratypes for A. g. gomerensis (locality of deposition unknown) are 2♂♂ dedicated by WARNCKE (1968) as paratypes of A. wollastoni acuta for La Gomera (El Cedro, ca. 1000 m, 23.03.1950, leg. Lindberg).

D i a g n o s t i c q u a l i t a t i v e f e a t u r e s: Inadditiontothemorphological characteristics that are typical for A. gomerensis , A. g. gomerensis is characterised by the following specific features:

F e m a l e: Colour. Mesosoma: femur, tibia, and basitarsus black (dark brown) similar to A. g. palmae nov. ssp. but in A. g. palmae nov. ssp. there are also few specimens (10%) with partly reddish colour; mediotarsi reddish-brown similar to A. palmae nov. ssp. but in A. g. gomerensis there are also some specimens (14%) with brown colour. Metasoma: T1- 4 black with black to dark reddish-brown depression zone (71%) but there are also specimens (29%) with dark reddish-brown depression zone; T5 depression zone reddishbrown.

Pubescence. Mesosoma: tibial scopa dorsally with brownish hairs (A. g. palmae nov. ssp. with slight brownish hairs), dorsobasal with reddish-brown hairs, tibial scopa ventral with yellowish-white hairs ( Fig. 15f ).

Structure. Head: clypeus without impunctate line ( Fig. 15b ) in contrast to A. g. palma e nov. ssp. with fragmented impunctate line (71%) or without impunctate line (29%); shallow more or less densely punctured (PD 28 μm, distance 14-56[70] μm), labrum process liguliform-trapezoidal (56%) ( Fig. 15b ) or trapezoidal (44%) in contrast to A. g. palmae nov. ssp. triangular , rounded on the top, laterally on the top more or less oblique. Mesosoma: scattered punctured with very shallow punctures, especially in front (PD =14 μm) ( Fig. 15c ).

M a l e: similar to female with following differences:

Colour. Head: flagellum brown (66%) or light brown (33%) ( Figs. 16a, c, d ) in contrast to A. g. palmae nov. ssp. light brown. Mesosoma: mediotarsi reddish-brown (66%) or brown (34%) in contrast to A. g. palmae nov.ssp. reddish-brown.

Pubescence. Metasoma: ST 8: long yellowish hairs at the end in contrast to A. g. palmae nov. ssp. with long white-yellowish hairs.

Structure. Head: vertex above the ocelli narrow, as wide as the ocellar diameter (80%) ( Fig. 16e ) or half of the diameter (20%) in contrast to A. g. palmae nov. ssp. with all specimens with the vertex as wide as the ocellar diameter; labrum process trapezoidal, emarginated, ends left and right side slightly thickened (80%) ( Fig. 17b ) or thickened (20%) in contrast to A. g. palmae nov. ssp. only slightly thickened.

D i s t r i b u t i o n: The distribution of this subspecies is with high probability limited to La Gomera ( Fig. 1c ). But there is one detection of A. g. gomerensis on El Hierro (section ‘ A. gomerensis , status’). The specimen (paratype in the OLML) was collected in El Hierro, Costa de Val verde, 28.04.1942, unknown collector). Despite intensive searching, no further specimens could be detected on El Hierro. According to HOHMANN et al. (1993) and our results, A. w. gomerensis is widely distributed in La Gomera. This subspecies covers altitudes between sea level (the northern and western coasts) up to 1200 m a.s.l.

F l i g h t p e r i o d: MarchandApril.

F u r t h e r s p e c i m e n s e x a m i n e d: 4♀♀ La Gomera, Ermita de Nuestra Señora de Las Nieves, 1073 m, 28°06'02.5''N, 17°12'13.6''W, 24.04.2016, leg. Kratochwil (ID-No KR-GO30 - KR-GO32, KR-GO35-KR-GO37, KR-GO42 ) ; 1♀ La Gomera, east of Temocodá , 1255 m, 28°06'40.4''N, 17°15'44.3''W, 24.04.2016, leg. Kratochwil (ID-No KR-GO34 ) ; 6♀♀ La Gomera, east of Temocodá , 1255 m, 28°06'40.4''N, 17°15'44.3''W, 24.04.2016, leg. Kratochwil (ID-No KR- GO34-1, KR-GO38-KR-GO41, KR-GO43 ) ; 3♀♀ La Gomera, Las Rosas , 626 m, 28°11'21.5''N, 17°13'15.3''W, 25.04.2016, leg. Kratochwil (ID-No KR-GO44 - KR-GO46 ) ; 6♀♀ La Gomera, northeast of Ermita de Nuestra Señora de Las Nieves, 615 m, 28°07'56.7''N, 17°12'11.7''W, 26.04.16, 25.04.2016, leg. Kratochwil (ID-No KR-GO47 - KR-GO52 ) ; 4♀♀ La Gomera, Ermita de Nuestra Señora de Las Nieves, 1073 m, 28°06'02.5''N, 17°12'13.6''W, 24.04.2016, leg. Kratochwil (ID-No KR-GO53 - KR-GO56 ) .

Figures

Fig 1: (a) Topographical location (with °North and °West of latitude and longitude) of the Azores, the Archipelago of Madeira, the Selvagens Islands, the Canary Islands and Cape Verde. (b) Islands of the Madeira Archipelago. (c) The Western Canary Islands and four areas of Tenerife (Anaga, Teno, Las Cañadas/Teide and the Dorsal Rift region). Modified according to AGUIAR & KARSHOLT (2006).

Fig. 15: Type specimen (holotype, female) of Andrena gomerensis WARNCKE, 1993 (OLML Nr. 6952779; ZOBODAT, 05.02.2020; biogeographical data set). (a) upper face with rugulae. (b) clypeus. (c) mesonotum. (d) propodeum. (e) abdomen. (f) scopa.

Fig. 16: Type specimen (paratype, male) of Andrena gomerensis WARNCKE, 1993 (OLML Nr. 6952778; ZOBODAT, 05.02.2020; biogeographical data set). (a) habitus dorsal. (b) type label. (c) head frontal. (d) head dorsal. (e) mesosoma. (f) metasoma.

Fig. 17: Type specimen (paratype, male) of Andrena gomerensis WARNCKE, 1993 (OLML Nr. 6952778; ZOBODAT, 05.02.2020; biogeographical data set). a: upper face with rugulae, b: clypeus, c: mesonotum, d: propodeum.