Hemerodromia alphalutea , Plant, Adrian R., 2015

Plant, Adrian R., 2015, Diversity of Hemerodromia Meigen, 1822 (Diptera: Empididae) in Thailand, the tip of a tropical iceberg?, Zootaxa 4039 (1), pp. 1-56: 10-11

publication ID


publication LSID


persistent identifier


treatment provided by


scientific name

Hemerodromia alphalutea

sp. nov.

Hemerodromia alphalutea  sp. nov.

( Figs 11 –13View FIGURES 11 – 13, 74View FIGURES 73 – 81)

Diagnosis. A species with scutum and scutellum yellow with mediotergite, laterotergite and metanepisternum on ventral part black. Sct present. Cell bm+dm long. Characteristically elongate pv spines on F 1. Male terminalia with complex cercus and epandrium with narrow dorsoapical process bearing long setae.

Description. Male: Body length 2.5 mm; wing length 2.3 mm. Head. Black, dusted greyish; antenna and mouthparts pale whitish yellow, all setae pale; one pair reclinate ocl; postocular setae uniseriate, becoming 2–3 serial below, merging above with 1–2 pairs small vtl hardly distinguishable from fine setulae on vertex and upper occiput; frontal setulae apparently absent. Antenna with postpedicel ~ 1.8 –2.0X long as wide, a few minute outstanding setulae subapically, stylus slightly shorter than postpedicel; scape with distinct fine dorsal seta. Thorax. Dusted greyish. Ground colour yellow with mediotergite, laterotergite and metanepisternum on ventral part black; scutum with vague indications of lateral stripes on posterior 0.4–0.5; prothoracic ‘collar’ yellow, narrowly blackish medially with suture between it and anterior margin of scutum dark; scutellum yellow, sutures at anterior and lateral margins dark. All setae yellowish; minute apart from a pair of distinct sct and one npl. Legs. Whitish yellow. C 1 ~ 1.2 X longer than distance between C 1 and C 2, all setulae minute excepting a few slightly longer apicals and a posterior ciliation of fine proclinate hairs ~ 0.3 X long as limb is deep. F 1 ~ 1.2 X long as C 1, ~ 6 X long as wide, evenly inflated, slightly constricted on proximal 0.2; femoral formula ~ 5– 6 / 17–19 / 16–19 + 1 / 5–6, denticles black, rows converging apically; spines yellow, pv row with ~ 3 spines between 0.2 and 0.7 obviously longer than limb is deep, av row shorter, becoming stronger proximally. T 1 ~ 0.8 X long as F 1, evenly curved, ventral face shallowly concave; with one row of ~ 17 sharply pointed spinose setae ventrally; a very small yellow ventroapical spur with distinct erect ventroapical blackish seta arising from near its base; a short ciliation of distinct proclinate hairs dorsally on distal 0.2. Mid and hind legs slender with only small pale setulae, somewhat longer dorsally on distal 0.3 of T 3. Wing. Membrane slightly tinged brownish by rather even covering of greyish brown microtrichia but hyaline basally. Veins brownish yellow to brown but whitish near base; cell bm+dm distally and veins R 2 + 3 and R 4 + 5 basally somewhat darker. Marginal setulae pale. R 2 + 3 linear on proximal 0.5, gently curved beyond, joining C ~ 0.6 X distance between end of R 1 and R 4; length of C between ends of R 2 + 3 and R 4 ~ 1.7 X long as R 4; R 4 almost linear, very faintly curved (slightly convex in second submarginal cell, R 4); R 5 ~ 2.0– 2.2 X long as R 4, almost linear; R 4 + 5 fork distal to M 1 + 2 fork by ~ 0.8–1.1 X length of R 4. M 1 almost linear beyond base, convergent distally with R 5; cell bm+dm moderately long, ending distinctly beyond tip of R 1. Halter yellowish white. Abdomen. Yellow with tergites 2–6 black. Terminalia. Black. Cercus ( Figs. 11, 12View FIGURES 11 – 13) of complex structure; extending slightly beyond epandrium; distally spatulate in lateral view with strong apical setae; a dorsally projected semitransparent subovate process on inner margin near base, bearing close ciliation of fine even-sized hairs at margin; immediately distal to this and originating from beneath it, a small inwardly directed club-shaped process bearing mostly lateroventrally and anteroventrally directed short spines; slightly distal to this a short rod-like process, inwardly directed, with 2 strong bristles apically. Epandrium rather rhomboid with distinct apical setae; a small narrow irregularly shaped lobe dorsoapically bearing long setae ( Fig. 11View FIGURES 11 – 13). Hypandrium somewhat bluntly pointed apically in ventral view ( Fig. 13View FIGURES 11 – 13); divided ventrobasally, fused ventroapically. Female: Unknown.

Type material. HOLOTYPE ♂, THAILAND: Chiang Mai Province, Doi Phahompok, Doi Phaluang, 1,449 m, 20 ° 1.06 'N, 99 ° 9.581 'E, 28.xi.– 5.xii. 2007, MT, P. Wongchai [T 6211]( QSBG). PARATYPES: 1 ♂, same data as holotype, ( QSBG); 1 ♂, Kamphaeng Phet Province, Mae Wong NP, 1,306 m, 16 ° 5.212 'N, 99 ° 6.576 'E, 12– 19.xi. 2007, C. Piluek & A. Inpuang ( NMWC).

Etymology. The specific epithet is a concatenation of the Latin luteus and the first letter of the Greek alphabet (alpha). It is used as a noun in apposition.

Remarks. This species may be separated from other predominantly yellow species by its characteristically elongate pv spines on F 1 and male terminalia with complex cercus and epandrium with narrow dorsoapical process bearing long setae. It is only known from hill evergreen forests at 1,306–1,449 m in Khamphaeng Phet and Chiang Mai provinces in northern Thailand ( Fig. 74View FIGURES 73 – 81). All specimens were captured in the cool dry season during November and December.


National Museum of Wales