Hemerodromia deltalutea , Plant, Adrian R., 2015

Plant, Adrian R., 2015, Diversity of Hemerodromia Meigen, 1822 (Diptera: Empididae) in Thailand, the tip of a tropical iceberg?, Zootaxa 4039 (1), pp. 1-56: 20-21

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Hemerodromia deltalutea

sp. nov.

Hemerodromia deltalutea  sp. nov.

( Figs 26 –28View FIGURES 26 – 28, 80View FIGURES 73 – 81)

Diagnosis. A yellow species with mediotergite obscured brownish and sutures at anterior margin of scutum, between lateral margin of scutum and fused anepisternum + katepisternum, at anterior and lateral margins of scutellum, at anterior margin of laterotergite and immediately anterior of posterior spiracle, very narrowly black. Sct present. Cell bm+dm long. Epandrium strongly concave on dorsal margin and surstylus with a ‘brush’ of very long black bristles apically.

Description. Male: Body length 2.8 mm; wing length 2.2 mm. Head. Black, dusted greyish, frons paler reddish black becoming paler still near base of antennae; antenna and mouthparts pale whitish yellow, all setae pale; one pair reclinate ocl; postocular setae uniseriate, becoming biserial below, merging above with 2–3 pairs small vtl hardly distinguishable from fine setulae on vertex and upper occiput; frontal setulae apparently absent. Antenna with postpedicel ~2.0X long as wide, a few minute outstanding setulae subapically, stylus slightly shorter than postpedicel; scape with distinct fine dorsal seta. Thorax. Dusted greyish. Ground colour yellow with mediotergite variably obscured brownish; sutures between prothoracic ‘collar’ and anterior margin of scutum, between lateral margin of scutum and fused anepisternum + katepisternum, at anterior and lateral margins of scutellum, at anterior margin of laterotergite and immediately anterior of posterior spiracle, very narrowly black. All setae yellowish; minute apart from a pair of distinct sct and one smaller npl. Legs. Whitish yellow. C 1 ~ 1.2 X longer than distance between C 1 and C 2, all setulae minute excepting a few slightly longer apicals and a posterior ciliation of fine proclinate hairs ~ 0.2 X long as limb is deep. F 1 ~ 1.2 X long as C 1, ~ 6 X long as wide, evenly inflated, slightly constricted on proximal 0.2– 0.3; femoral formula ~ 6– 7 / 17–20 / 17–20 + 2 / 6–7, denticles black, rows converging apically; spines yellow, pv and av rows similarly long, becoming somewhat longer proximally. T 1 ~ 0.8 X long as F 1, evenly curved, ventral face shallowly concave; with one row of ~ 17 sharply pointed spinose setae ventrally; a very small yellow ventroapical spur with distinct ventroapical erect black or yellowish black spinose seta arising from near its base; a short ciliation of distinct proclinate hairs dorsally on distal 0.2. Mid and hind legs slender with only small pale setulae, somewhat longer dorsally on distal 0.3 of T 3. Wing. Membrane slightly darkened by rather even covering of greyish brown microtrichia but hyaline basally. Veins brownish yellow, somewhat darker about base of R 2 + 3, R 4 + 5 and distal part of cell bm+dm but pale yellowish near base. Marginal setulae pale. R 2 + 3 linear on proximal 0.5, gently curved beyond, joining C ~ 0.7 X distance between end of R 1 and R 4; length of C between ends of R 2 + 3 and R 4 ~ 1.5–1.8 X long as R 4; R 4 almost linear, only very vaguely Sshaped; R 5 ~ 2.0– 2.2 X long as R 4, almost linear; R 4 + 5 fork distal to M 1 + 2 fork by ~ 1.0– 1.2 X length of R 4. M 1 almost linear beyond base, convergent distally with R 5; cell bm+dm moderately long, ending distinctly beyond tip of R 1. Halter yellowish white. Abdomen. Yellow with tergites 4–6 entirely, 2–3 broadly on at least distal ~ 0.5 and 1 very narrowly on posterior margin black. Terminalia. Black or brownish-black with cercus paler, yellowish brown. Cercus ( Figs 26, 27View FIGURES 26 – 28) of complex structure; about as long as epandrium; subtriangular in dorsal view. Apical lobe incurved; broad, sub-stellate or somewhat club-shaped in lateral view. Median lobe less strongly sclerotized, incurved, bearing a few fine bristly setae apically. Epandrium ( Fig. 26View FIGURES 26 – 28) with dorsal margin strongly concave distally in lateral view; a short bluntly pointed dorsoapical projection appearing sharper and longer in posterior view. Surstylus present, emerging slightly beyond end of epandrium; with dense ciliation of black flattened specialised setae apically. Hypandrium rather triangular in ventral view ( Fig. 28View FIGURES 26 – 28) with apex broadly rounded; very long black setae apically, becoming shorter and yellowish proximally, bare basally; fused ventroapically, divided ventrobasally. Female: Unknown.

Type material. HOLOTYPE ♂, THAILAND: Kamphaeng Phet Province, Mae Wong NP, Chong Yen, 1,306 m, 16 ° 5.212 'N, 99 ° 6.576 'E, 10–17.xii. 2007, MT, C. Piluek & A. Inpuang [T 6302]( QSBG). PARATYPES: 5 ♂, same data as holotype, 10–17.xii.2007, 24– 31.xii.2007, 7– 14.i. 2008 ( QSBG & NMWC).

Additional material. Three female Hemerodromia  specimens collected at the same locality as the holotype male on 24–31.xii. 2007 and 7–14.i. 2008 ( QSBG) may belong to this species but could not be confidently be assigned to it.

Etymology. The specific epithet is a concatenation of the Latin luteus and the fourth letter of the Greek alphabet (delta). It is used as a noun in apposition.

Remarks. The ‘brush’ of very long black bristles on the surstylus is a character that readily distinguishes Hemerodromia deltalutea  sp. nov. from other predominantly yellow species of the genus in Thailand. The species is only known from a single locality in hill evergreen forest at 1,306 m in Kamphaeng Phet Province in northern Thailand ( Fig. 80View FIGURES 73 – 81) during December and January in the cool dry season.


National Museum of Wales