Hemerodromia epsilutea , Plant, Adrian R., 2015

Plant, Adrian R., 2015, Diversity of Hemerodromia Meigen, 1822 (Diptera: Empididae) in Thailand, the tip of a tropical iceberg?, Zootaxa 4039 (1), pp. 1-56: 24-26

publication ID


publication LSID


persistent identifier


treatment provided by


scientific name

Hemerodromia epsilutea

sp. nov.

Hemerodromia epsilutea  sp. nov.

( Figs 34 –36View FIGURES 34 – 36, 83View FIGURES 82 – 90)

Diagnosis. A species with thorax entirely yellow with only sutures in front of root of halter and about laterotergite blackish. Sct present. Cell bm+dm long. Abdomen with tergite 2 blackish. Male terminalia with hypandrium and epandrium clear yellow, surstylus broadly pointed emerging beyond tip of epandrium and bearing strong setae apically.

Description. Male: Body length 1.9 mm; wing length 1.8 mm. Head. Black, dusted greyish, frons slightly paler; antenna and mouthparts pale whitish yellow, all setae pale; one pair small reclinate ocl; postocular setae very small, becoming somewhat longer below, merging above with 1–2 pairs small vtl and a few fine hairs on vertex; frontal setulae apparently absent. Antenna with postpedicel ~ 2.5 X long as wide, a few minute outstanding setulae subapically; stylus somewhat shorter than postpedicel; scape with distinct fine dorsal seta. Thorax. Dusted greyish. Ground colour clear yellow including mediotergite and prothoracic ‘collar’; only sutures in front of root of halter and about laterotergite blackish; anterior spiracle narrowly and faintly emarginated above with diagonal dark mark. All setae yellowish; minute apart from a pair of distinct sct and one smaller npl. Legs. Pale yellow. C 1 ~ 1.1–1.2 X longer than distance between C 1 and C 2, all setulae minute. F 1 ~ 1.2 X long as C 1, ~ 6 X long as wide, evenly inflated, hardly constricted on proximal 0.3; femoral formula ~ 6 / 15 / 16 + 1 / 6, denticles black, rows converging apically; spines yellow, pv and av rows similarly very short and hair-like, becoming somewhat longer proximally. T 1 ~ 0.6–0.7 X long as F 1, evenly curved, ventral face shallowly concave; with row of ~ 16 sharply pointed spinose setae ventrally (in the holotype T 1 is tightly reflexed against F 1 and it is uncertain if it bears 1 or 2 rows of setae); ventroapical spur small with distinct ventroapical erect black spinose seta arising from near its base. Mid and hind legs slender with only small pale setulae. Wing. Membrane faintly darkened by rather even covering of brownish yellow microtrichia but hyaline basally. Veins brownish yellow, somewhat darker about junction of R 1 and C, base of R 2 + 3, R 4 + 5 and cell bm+dm, but paler near base. Marginal setulae pale. R 2 + 3 almost linear, joining C ~ 0.6–0.7 X distance between end of R 1 and R 4; length of C between ends of R 2 + 3 and R 4 ~ 1.8 –2.0X long as R 4; R 4 slightly Sshaped, angle with R 5 at extreme base ~ 85 °, becoming ~ 70–80 ° thereafter, joining C at ~ 85 °. R 5 ~ 2.8 X long as R 4, slightly curved; R 4 + 5 fork distal to M 1 + 2 fork by ~ 1 X length of R 4. M 1 almost linear beyond base, convergent distally with R 5; cell bm+dm long, ending beyond tip of R 1. Halter yellowish white. Squama with margin greyish. Abdomen. Yellow with tergites 2–5 blackish; all setae yellow, short but slightly longer on distal segments, especially sternites 6 and 7. Terminalia. Epandrium and hypandrium clear yellow, cercus and surstylus black. Cercus distinctly angulate in dorsal view ( Fig. 34View FIGURES 34 – 36), distal lobe incurved bearing numerous strong setae ( Fig. 35View FIGURES 34 – 36). Epandrium sub-trapezoid, without strong setae on external face. Surstylus present, broad bluntly pointed apically with strong apical setae, partly fused with and emerging beyond epandrial lamella. Hypandrium narrow, with distinct setae distally. Phallus with sharply pointed apical processes ( Fig. 36View FIGURES 34 – 36). Female: Unknown.

Type material. HOLOTYPE ♂, THAILAND: Chiang Mai Province, Doi Inthanon NP, Wachirathan trap 1, 662 m, 18.5408 °N, 98.6001 °E, 2–29.v. 2014, MT, W. Srisuka R. Sawkord T. Somboonchai & S. Suriya [QSBG- 2014 - 134]( QSBG).

Etymology. The specific epithet is a concatenation of the Latin luteus and the fifth letter of the Greek alphabet (epsilon). It is used as a noun in apposition.

Remarks. The combination of male terminalia clear yellow and abdomen yellow with tergite 2 blackish serves to distinguish H. epsilutea  sp. nov. from other yellow species of Hemerodromia  occurring in Thailand although confirmation of identity requires more detailed examination of the male terminalia. The species is only known from a single lowland (662 m) site in dry evergreen forest during May in Chiang Mai Province, northern Thailand ( Fig. 83View FIGURES 82 – 90).