Lanzatus somalilandus Kovařík et Lowe, 2016

Kovařík, František, Lowe, Graeme & Šťáhlavský, František, 2016, Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part IX. Lanzatus, Orthochirus, and Somalicharmus (Buthidae), with description of Lanzatus somalilandus sp. n. and Orthochirus afar sp. n., Euscorpius 232, pp. 1-38 : 5-9

Lanzatus somalilandus Kovařík et Lowe , sp. n.

( Figs. 1–4, 7–10, 13–28, 30–31, 35–40, 123–124, 155, Table 1) http://zoobank.org/urn:lsid:zoobank.org:act:22B29B

26-A917-4D0E-BF1F-6D5A1EEB7992

Lanzatus somalicus: Kovařík et al., 2013a: 18 .

TYPE LOCALITY AND TYPE REPOSITORY. Somaliland , between Sheikh and Laas Caanood, 09°36'40.1"N 45°29' 35.7"E, 1089 m a.s.l. , FKCP.

TYPE MATERIAL. Somaliland , between Sheikh and Laas Caanood, 09°36'40.1"N 45°29'35.7"E, 1089 m a.s.l. (Locality No. 11SL, Fig. 40 and fig. 71 in Kovařík et al., 2013a: 17), 10.VII.2011, 2♂ (holotype, Figs. 3–4, 9–10, 28, 30–31 and paratype) 1♀ (paratype, Figs. 1–2, 7–8, 13–27, 35–37, 39, 123–124), leg. F. Kovařík , FKCP.

ETYMOLOGY. Named after the country of occurrence.

DIAGNOSIS. Total length 17.9 (male)– 26 mm (female). Base color uniformly yellowish orange with black only around the eyes. Fifth metasomal segment slightly marbled in grayish black. Pectine teeth number 18 in both sexes. Pedipalps bear inconspicuous smooth carinae. Pedipalp chela smooth and narrow. Sternites smooth, without carinae and granules. Tergites I–VI finely granulated with one carina present or indicated. Metasomal segments smooth, without carinae, with rounded edges. Dorsal surfaces of metasomal segments I–IV bear an inconspicuous longitudinal groove. Metasomal segments III–V smooth (female) or densely and very finely granulated (males). Telson elongated, smooth, with aculear ring slightly indicated in male. Aculeus strongly curved.

DESCRIPTION. The adults are 17.9 (male)– 26 mm (female) long. The habitus is shown in Figs. 1–4. For position and distribution of trichobothria of pedipalps see Figs. 17–20 and 22. Sexual dimorphism: adult male integument matte, female glossy ( Figs. 38–39), telson narrower in male ( Fig. 28) than in female ( Fig. 27), with aculear ring indicated.

Coloration ( Figs. 38–39). The base color is uniformly yellowish orange with black only around the eyes, tergites darker. Tarsomeres of legs and telson white. Fifth metasomal segment slightly marbled in grayish black.

Carapace and mesosoma ( Figs. 7–10). Entire carapace is finely granulated, with a glossy area only between the posterior and median eyes in the female. Carinae absent. Anterior margin of carapace almost straight. Tergites finely granulated, with one median carina more indicated in female. Pectinal tooth count 18 in both sexes. Pectine marginal tips extend to half of the fourth sternite in the female, to end of the fourth sternite in the male. Pectines with 3 marginal lamellae and 8–9 middle lamellae. All lamellae and fulcra bear numerous pale setae. All sternites smooth.

Metasoma and telson ( Figs. 27–28, 30–31, 35–37). Metasomal segments smooth and hirsute, without carinae, with rounded edges. Dorsal surface of the first to fourth metasomal segments bearing an inconspicuous longitudinal groove. Third to fifth metasomal segments smooth (female) or densely and very finely granulated (males). Telson elongated, smooth, with aculear ring slightly indicated in male. Telson length/ width ratio 3.94 in male, 3.61 in female. Aculeus strongly curved.

Chelicerae ( Figs. 13–14, 123–124). Manus 1.4 times longer than wide, dorsal surface smooth with strong anterior marginal and dorsointernal carinae, bearing 3 long pale setae, 2 on anterior marginal carina and 1 submarginal, and 1 long dark macroseta on mid-dorsointernal carina. Dorsal surface of movable finger smooth, with 2 pale microsetae. Dorsal surface of fixed finger with sparse setae with fluorescent bases. Ventral surfaces of manus, fixed and movable fingers bearing numerous medium to long setae with fluorescent bases ( Fig. 14), setae on ventral manus well spaced, not very dense, widely distributed. Fingers robust, with typical buthid dentition (Vachon, 1963). Dorsal margin of movable finger armed with 5 teeth: dorsal distal tine, subdistal, median and apparently 2 small basal teeth fused in bicusp. Ventral margin of movable finger with 3 teeth: ventral distal tine, median and basal teeth. Fixed finger margin with 4 teeth: distal tine, subdistal, median and basal teeth fused into bicusp. Ventral surface of fixed finger armed with a single tooth ( Fig. 124).

Pedipalps ( Figs. 15–22). Pedipalps smooth, sparsely hirsute, with inconspicuous smooth carinae. Pedipalp chela narrow, chela length/ width ratio 5.23 in male, 5.28 in female. Dentate margin of pedipalp chela with distinct granules divided into 7 diagonal rows (including apical row) on movable finger, 6 diagonal rows on fixed finger (of which five have an accessory external granule). Terminal and subterminal granules present.

Legs ( Figs. 23–26). Legs I–IV with tibial spurs absent, retrolateral and prolateral pedal spurs present. All legs smooth, without distinct carinae. Tarsomeres bear two rows of macrosetae on the ventral surface, and additional macrosetae on other surfaces. Tarsomeres of legs I–III with 5 to 9 setae.

Measurements. See Table 1.

A FFINITIES. Lanzatus somalilandus sp. n. and L. somalicus are differentiated mainly by separate areas of distribution ( Fig. 155). They can be separated by morphology of the telson, which is narrower with aculear ring only indicated in L. somalilandus sp. n. ( Fig. 28), vs. broader with ring well developed in a male of L. somalicus ( Fig. 29). The telson length/ width ratio is 3.94 in male and 3.61 in female of L. somalilandus sp. n., vs. 3.45 in male paratype of L. somalicus . The aculeus is strongly curved in L. somalilandus sp. n. ( Figs. 27–28) vs. slightly curved in L. somalicus ( Fig. 29). L. somalicus has 19–24 pectine teeth in males (19– 20 in material examined here, 24 in type of Sabinebuthus elegans ), vs. 18 teeth in all three types including two males of L. somalilandus sp. n.

C OMMENTS ON LOCALITIES AND LIFE STRATEGY. On 10 July 2011 the first author, Tomáš Mazuch and Pavel Novák stopped at the locality 11SL (c.f. Fig. 40 and fig. 71 in Kovařík et al., 2013a: 17; a sizeable area of windblown sands with sparse shrubs) about one hour after sunset and persuaded the soldiers/ protectors to allow collecting near a road by UV detection at night for at least 15 minutes, a period which yielded all three types of L. somalilandus sp. n., found running on the sand near the shrubs, and a juvenile paratype of Gint dabakalo Kovařík et Mazuch, 2015 .