Stuckenbergomyia secunda (Smith, 1969)

Sinclair, Bradley J., 2019, Revision of the southern African genus Stuckenbergomyia Smith, 1971 (Diptera, Empidoidea) and proposal of a new subfamily, African Invertebrates 60 (1), pp. 133-145 : 133

Stuckenbergomyia secunda (Smith, 1969) Figs 8-12, 13-14, 19, 22

Stuckenbergia secunda Smith, 1969: 128. Type locality: Pietermaritzburg, KwaZulu-Natal, South Africa.

Stuckenbergomyia secunda : Smith, 1980: 436 (catalogue).

Diagnosis.

This species is characterised by the faint brown vittae beneath acrostichals and dorsocentrals; broad wings and 4 pairs of long scutellar setae; male terminalia: symmetrical, held at apex of abdomen, surstyli broadly articulated lobes.

Redescription.

Head: Holoptic with enlarged facets on upper two-thirds. Antenna dark brown; postpedicel length ca. 4 × basal width; stylus cylindrical, about as long as basal width of postpedicel; apical mechanoreceptor one-third length of stylus.

Thorax: Mostly dark greyish pruinescence. Scutum with brownish vittae below acrostichals and dorsocentral rows (viewed anteriorly); anterior margin of postpronotal lobe shiny, lacking pruinescence. Acrostichals 4-serial anteriorly, biserial anterior to prescutellar depression; dorsocentral setae biserial and short anteriorly, longer, darker and uniserial at prescutellar depression; postpronotal setae numerous, long and pale; numerous pale presutural and postsutural supra-alar setae; 4 pale notopleural setae; 1 pale postalar seta; 4 pairs of pale scutellar setae, longer than length of scutellum. Proepisternum with several long, pale setae.

Legs: Mostly brown, with apex of femora and base of tibiae yellowish (Fig. 10).

Wing (2.7-3 mm) (Fig. 11): Broad, anal lobe nearly right-angled; alula well-developed. Costal margin at humeral crossvein expanded, costal cell broad; costa strongly reduced beyond R4+5; spur vein arising from cell cua. Halter knob pale.

Abdomen: Slightly paler than thorax; setae pale; sternites paler than tergites; tergite 8 narrow medially, expanded laterally, bearing pair of setae.

Male terminalia (Figs 13, 14): Symmetrical, held at apex of abdomen, directed dorsally, not arched over abdomen; rotated 45° to right. Cercus quadrate, thinly sclerotised, weakly pigmented, clothed in slender setae; hypoproct broad with fine setae on posterior margin, arched medially. Epandrial lamellae very narrowly joined; surstylus broadly articulated with sickle-shaped, arched posterior process. Hypandrium with pair of setae; postgonite consisting of 2 pairs of sickle-shaped, slender, processes; ventral apodeme broad, plate-like. Phallus and ejaculatory apodeme not examined.

Female.

Unknown.

Type locality.

SOUTH AFRICA, Pietermaritzburg [29°36'S, 30°22'E] (Fig. 22).

Type material examined.

Holotype ♂, labelled (Fig. 8): "Pietermaritzburg / South Africa / B. & P. Stuckenberg"; "grassland/ 31.5.62 [underside of label]"; "HOLOTYPE ♂ / Stuckenbergia / secunda / K.G.V. SMITH"; " NMSA-DIP-09434" (NMSA) [dissected, genitalia mounted on microslide beneath specimen] (Fig. 9).

Additional material examined.

SOUTH AFRICA: 2♂ [NMSA-DIP 092039, NMSA-DIP 092040], KwaZulu-Natal: Louwsberg, Sanyati Farm, 1090 m elev., 27°34'90"S, 31°17.9'E, 11-13.vii.2006, YPT, M. Mostovski, leg (NMSA).