Eoanaphes Huber

Huber, John T. & Greenwalt, Dale, 2011, Compression fossil Mymaridae (Hymenoptera) from Kishenehn oil shales, with description of two new genera and review of Tertiary amber genera, ZooKeys 130, pp. 473-494: 473

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scientific name

Eoanaphes Huber

gen. n.

Eoanaphes Huber  ZBK  gen. n. Figs 17, 18


Female. Body about 700. Head. About as wide as high (measured in frontal view). Malar space long, almost eye height. Antenna. Funicle 6-segmented, with fl1 very short and remaining segments longer than wide; clava 3-segmented with the claval sutures almost perpendicular to claval length. Wings. Fore wing wide, slightly asymmetrical, with somewhat truncate apex; marginal fringe much shorter than fore wing width. Venation almost 1/3 fore wing length, with marginal vein fairly short, about as long as stigmal vein. Hind wing narrow; marginal fringe much longer than wing width and along posterior margin extending to base of membrane.


About 0.75 x gaster length. Mesoscutum length shorter than scutellum. Metanotum much shorter than scutellum, apparently with strap-like dorsellum. Metasoma. Apparently slightly constricted at base, probably with fairly wide, short petiole. Gastral terga similar in length. Ovipositor moderately short, apparently arising near base of gaster and its apex not exserted beyond gastral apex.

Type species.

Eoanaphes stethynioides  Huber, sp. n.

Derivation of generic name.

The name is based on the extant genus Anaphes  , which is also known from one extinct species in Baltic amber. The gender is masculine.


Eoanaphes  appears to be related to two genera, Anaphes  and Stethynium  . Four features suggest Anaphes  : 1) in lateral view there is a distinct constriction dorsally between the mesosoma and metasoma, indicating that the propodeum slopes strongly down relative to the horizontal scutellum and dorsellum (in Stethynium  , the entire dorsal margin of the mesosoma is horizontal and in line with the base of the metasoma, without a depression between the two parts); 2) the very short fl1; 3) the apparent absence of a distinct rounded lobe on the posterior margin of the fore wing opposite the marginal + stigmal veins (one wing appears to have a rounded lobe but this is due to the membrane being partly folded over on itself); 4) the wing surface behind the marginal vein seems to be bare except for a few setae behind the stigmal vein and a faint suggestion of a hair line separating medial from marginal space. Two features suggest Stethynium  : 1)the clearly 3-segmened clava; 2) the strap-like dorsellum (rhomboidal in Anaphes  ).