Haplodrassus rhodanicus ( Simon, 1914 )

Bosmans, Robert, Kherbouche-Abrous, Ourida, Benhalima, Souâd & Hervé, Christophe, 2018, The genus Haplodrassus Chamberlin, 1922 in the Mediterranean and the Maghreb in particular (Araneae: Gnaphosidae), Zootaxa 4451 (1), pp. 1-67: 29-33

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Haplodrassus rhodanicus ( Simon, 1914 )

comb. n.

Haplodrassus rhodanicus ( Simon, 1914)  , comb. n.

Figs 118‒125View FIGURES 118–125, 136‒139View FIGURES 136–144, map 3

Drassodes rhodanicus Simon, 1914: 122  , 134, 208 (♂).

Types. Male syntypes from France, Ardèche, Bourg St.-Andéol (N44°22’32” E4°38’33”) and Gard, Pont-St-Esprit (44°15’17” E4°38’50”); not examined, missing in MNHN where the type material normally should be hosted.

Remark. The male of this species is illustrated for the first time, and the female is described for the first time.

Diagnosis. This species is easily recognised in the male by the narrow, truncate tibial apophysis ( Figs 120View FIGURES 118–125, 137View FIGURES 136–144) and the narrow embolic apophysis and embolus ( Figs 122View FIGURES 118–125, 136View FIGURES 136–144). The female is recognized by the elongate epigyne and is clearly related to Haplodrassus longivulva  sp. n., from which it differs by the straight lateral margins of the fovea ( Figs 124View FIGURES 118–125, 138View FIGURES 136–144).

Description. Measurements: Male (n=12): Total length 2.3‒5.2; carapace 1.20‒2.12 long, 0.96‒1.73 wide. Female (n=3): Total length 2.9‒3.4; carapace 1.14‒1.33 long, 0.91‒0.98 wide. Colour: Carapace yellowish brown with darkened eye region; chelicerae brown to dark brown; sternum yellowish brown with brown margins; legs yellowish brown; abdomen grey, anterior half often with three pairs of drop-like spots. Spinulation: General pattern, but all metatarsi spineless and lateral spines on Fe III ‒IV missing.

Male palp ( Figs 120‒123View FIGURES 118–125, 136‒137View FIGURES 136–144): Tibia much longer than wide (ratio 2.6), apophysis shorter than tibia’s diameter (ratio 0.6), narrow from its base, terminally truncate with blunt tip ( Figs 120View FIGURES 118–125, 137View FIGURES 136–144); embolic apophyis small and elongated, terminally bluntly pointed; embolus narrow, with subterminal denticle ( Figs 122View FIGURES 118–125, 136View FIGURES 136–144).

Epigyne ( Figs 124‒125View FIGURES 118–125, 138‒139View FIGURES 136–144): Strongly elongated, nearly twice as wide as long, with rounded anterior hood; fovea rectangular, with straight margins ( Figs 124View FIGURES 118–125, 138View FIGURES 136–144); receptacles small and touching, with anterolaterally directed ducts ( Figs 125View FIGURES 118–125, 139View FIGURES 136–144).

Further material examined. AFRICA, TUNISIA, Jendouba, Hammam Bourguiba (N36°46'32” E8°36'35”), 350m, 1♂, stones in Quercus suber  forest, 9.V.2006, R. Bosmans leg. (CRB). EUROPE, PORTUGAL, Setubal, Alcochete NW (N38°45'37” W8°56’6”), 5m, 5♂, pitfalls in salt marsh, 14‒22.IV.2013, R. Bosmans leg. (CRB). SPAIN, Baleares, Mallorca, Sa Rapita W., Punta Negra (N39°21'22" E'2°54'43'), 10m, 1♀, stones in small lagoon near the sea, 5.IV.2003, R. Bosmans leg. (CRB), Málaga, Málaga (N36°43’12” W4°25’13”), 2♂ 1♀, as Drassus scenicus L. Koch  nomen nudum, E. Simon leg., Coll. Simon 2051 (MNHN AR9377), Palencia, Laguna de la Nava (N39°2’58” W3°41’47”), 670m, 2♂, 24.VI.1996, U. Stengele leg. (CRB), Valencia, Embalse de Cofrentes (N39°14'31” W1°3'37”), 375m, 2♂ 1♀, in Tamarix  and Phragmites  litter near the water, 3.IV.1996, R. Bosmans leg. (CRB). FRANCE, Aude, La Palme near Leucate (N42°58’53” E3°1’01”), 1♀, stones in salt marsh, 9.V.2015, J. Van Keer leg. (CJVK), Leucate, Chemin de Las Pichinos (N42°56’10” E3°1’8”), 4♂, pitfalls in salt marsh, 9‒26.V.2015 (CJVK), Bouches-du-Rhone, Petite Camargue (N43°30'0” E4°19'0”), 1♂ 1♀, 15.V.1989, P. Poot leg. (CRB), Haute-Corse: Etang de Biguglia near Borgo (N42°38’23” E9°26’39”), 15m, 1♀, pitfalls in Phragmites  stand bordering the lake, 28.V.1995, R. Bosmans leg. (CRB). ITALY, Sardinia, Medio Campidano, Montevecchio, Piccalinna (N39°33'24” E8°34'6”), 1♂, pitfalls in garrigue, 3.VI.2009, M. Verdinelli & A. Sassu leg. (MCSNB).

Ecology. The species was mostly collected in marshy areas: lagoons near the sea, inland salt marshes, along inside lakes and reservoirs. Males were found from April to June, females in April and May.

Distribution. Haplodrassus rhodanicus  was first described from the south of France ( Simon 1914) and later only cited from Croatia, Dubrovnik - Pobrezje ( Nikolic & Polenec 1981). The species now appears to have a much larger distribution area, with first records in Tunisia, Spain, Portugal and Italy. It was mostly collected in marshes (map 3).


Museum National d'Histoire Naturelle














Haplodrassus rhodanicus ( Simon, 1914 )

Bosmans, Robert, Kherbouche-Abrous, Ourida, Benhalima, Souâd & Hervé, Christophe 2018

Drassodes rhodanicus

Simon, 1914 : 122