Haplodrassus

Bosmans, Robert, Kherbouche-Abrous, Ourida, Benhalima, Souâd & Hervé, Christophe, 2018, The genus Haplodrassus Chamberlin, 1922 in the Mediterranean and the Maghreb in particular (Araneae: Gnaphosidae), Zootaxa 4451 (1), pp. 1-67: 7-8

publication ID

https://doi.org/10.11646/zootaxa.4451.1.1

publication LSID

lsid:zoobank.org:pub:5DCE18B3-9DBA-48F5-AEF7-90A50B984EA5

persistent identifier

http://treatment.plazi.org/id/DE5087A6-C013-B331-FF39-7404FA53FC3E

treatment provided by

Plazi

scientific name

Haplodrassus
status

 

Key to the Haplodrassus  species of the Maghreb region

MALES

1 Cymbium with abundant, strong spines (fig. 71)................................................... nigroscriptus 

- Cymbium with few, less strong spines (fig. 32).............................................................. 2

2(1) Embolic apophysis hooked (fig. 190)........................................................ triangularis  sp. n.

2 Embolic apophysis sigmoid or nearly straight ( Figs 4View FIGURES 1–9, 14View FIGURES 10–17, 35View FIGURES 30–40 …)................................................. 3

3(2) Embolic apophysis small, occupying only one third of tegulum ( Figs 122View FIGURES 118–125, 136View FIGURES 136–144); tibial apophysis slender, with narrow base ( Figs 120View FIGURES 118–125, 137View FIGURES 136–144)............................................................................... rhodanicus 

- Embolic apophysis large, occupying at least half of tegulum ( Figs 4View FIGURES 1–9, 14View FIGURES 10–17, 35View FIGURES 30–40 …); tibial apophysis voluminous, with broad base ( Figs 5View FIGURES 1–9, 12View FIGURES 10–17, 33View FIGURES 30–40 …)..................................................................................... 4

4(3) Embolic apophysis with dorsal keel ( Figs 158View FIGURES 154–161, 169View FIGURES 162–171)..................................................... signifer 

- Embolic apophysis without dorsal keel ( Figs 4View FIGURES 1–9, 14View FIGURES 10–17, 35View FIGURES 30–40 …)...................................................... 5

5(4) Embolic apophysis with 2 prolateral teeth ( Figs 14View FIGURES 10–17, 26View FIGURES 18–29, 93View FIGURES 87–97, 110View FIGURES 108–117)................................................. 6

- Embolic apophysis without ( Figs 4View FIGURES 1–9, 35View FIGURES 30–40, 103View FIGURES 98–107 …) or at most with 1 prolateral tooth ( Figs 62View FIGURES 58–66, 129View FIGURES 126–135)....................... 7

6(5) Embolic apophysis slender, elongated and with pointed tip ( Figs 93View FIGURES 87–97, 108View FIGURES 108–117).................................... omissus 

- Embolic apophysis compact, robust, with blunt tip ( Figs 14View FIGURES 10–17, 25View FIGURES 18–29)........................................ dalmatensis 

7(6) Embolic apophysis with one prolateral tooth ( Figs 62View FIGURES 58–66, 77View FIGURES 77–86, 129View FIGURES 126–135, 140View FIGURES 136–144)............................................. 8

- Embolic apophysis without prolateral tooth ( Figs 4View FIGURES 1–9, 35View FIGURES 30–40, 103View FIGURES 98–107 …)................................................. 9

8(7) Teeth on embolic apophysis equal, apophysis appearing bifid ( Figs 62View FIGURES 58–66, 79View FIGURES 77–86)............................... lyndae  sp. n.

- Teeth on embolic apophysis not equal (fig. 129, 142).................................................... rufipes 

9(8) Embolus with a strong retrolateral tooth at its base ( Figs 49, 51View FIGURES 49–57)....................................... dentifer  sp. n.

- Embolus without retrolateral tooth at base ( Figs 4View FIGURES 1–9, 45View FIGURES 41–48, 103View FIGURES 98–107 …)............................................ …….10

10(9) Tibial apophysis nearly symmetric ( Figs 100View FIGURES 98–107, 114View FIGURES 108–117).................................................. ovatus  sp. n.

- Tibial apophysis strongly asymmetric ( Figs 5View FIGURES 1–9, 43View FIGURES 41–48, 147View FIGURES 145–153, 186View FIGURES 181–189)................................................... 11

11(10) Tibial apophysis terminally strongly pointed ( Figs 186View FIGURES 181–189, 195View FIGURES 190–197).............................................. typhon 

- Tibial apophysis bluntly pointed ( Figs 5View FIGURES 1–9, 43View FIGURES 41–48, 147View FIGURES 145–153)........................................................... 12

12(11) Tibial apophysis gradually narrowing, with subterminal concavity ( Figs 43View FIGURES 41–48, 55View FIGURES 49–57)....................... longivulva  sp. n.

- Tibial apophysis not gradually narrowing, with basal concavity ( Figs 5View FIGURES 1–9, 147View FIGURES 145–153)..................................... 13

13(12) Embolic apophysis terminally with pointed nipple ( Figs 6View FIGURES 1–9, 20View FIGURES 18–29)........................................... crassipes 

- Embolic apophysis terminally blunt ( Figs 149View FIGURES 145–153, 164View FIGURES 162–171)............................................... securifer  sp. n. FEMALES

1 Tarsus and tibia of palp with more than 20 ventral spines (fig. 73)..................................... nigroscriptus 

- Tarsus of palp with less than 10 ventral spines (fig. 37........................................................ 2

2(1) Fovea triangular ( Figs 178View FIGURES 172–180, 192View FIGURES 190–197)............................................................ triangularis  sp. n.

- Fovea oval or rounded ( Figs 7View FIGURES 1–9, 16View FIGURES 10–17, 38View FIGURES 30–40 …)................................................................... 3

3(2) Epigyne at least twice as long as wide ( Figs 47View FIGURES 41–48, 64View FIGURES 58–66, 124View FIGURES 118–125)....................................................... 4

- Epigyne less elongate, at most 1.7 times as long as wide ( Figs 7View FIGURES 1–9, 16View FIGURES 10–17, 38View FIGURES 30–40).......................................... 6

4(3) Fovea narrow anteriorly, widening distinctly to the back ( Figs 64View FIGURES 58–66, 81View FIGURES 77–86)................................... lyndae  sp. n.

- Fovea more or less rectangular ( Figs 47View FIGURES 41–48, 124View FIGURES 118–125)............................................................... 5

5(4) Lateral margin of fovea with median angularity ( Figs 47View FIGURES 41–48, 56View FIGURES 49–57)...................................... longivulva  sp. n.

- Lateral margin of fovea with parallel margins ( Figs 124View FIGURES 118–125, 138View FIGURES 136–144)........................................... rhodanicus 

6(5) Fovea with postero-median septum ( Figs 16View FIGURES 10–17, 95View FIGURES 87–97)............................................................. 7

- Fovea without postero-median septum ( Figs 7View FIGURES 1–9, 38View FIGURES 30–40, 105View FIGURES 98–107)....................................................... 8

7(6) Fovea distinctly longer than wide ( Figs 17View FIGURES 10–17, 28View FIGURES 18–29)..................................................... dalmatensis 

- Fovea as long as wide ( Figs 95View FIGURES 87–97, 111View FIGURES 108–117)................................................................. omissus 

8(7) Foveal margin with distinct median tooth in lateral margin, dividing fovea into a trapezoid or rectangular anterior part and a rounded posterior part ( Figs 40View FIGURES 30–40, 132View FIGURES 126–135)...................................................................... 9

- Foveal margin without such tooth, fovea oval or rounded ( Figs 7View FIGURES 1–9, 105View FIGURES 98–107, 151View FIGURES 145–153, 160View FIGURES 154–161, 187View FIGURES 181–189).............................. 10

9(8) Areola anteriorly truncate ( Figs 132View FIGURES 126–135, 143View FIGURES 136–144)............................................................. rufipes 

- Areola anteriorly gradually narrowing ( Figs 40View FIGURES 30–40, 52View FIGURES 49–57)................................................ dentifer  sp. n.

10(9) Fovea longer than wide ( Figs 105View FIGURES 98–107, 187View FIGURES 181–189)................................................................... 11

- Fovea as long as wide ( Figs 7View FIGURES 1–9, 151View FIGURES 145–153, 160View FIGURES 154–161).................................................................. 12

11(10) Fovea oval, anterior margins converging ( Figs 105View FIGURES 98–107, 116View FIGURES 108–117).............................................. ovatus  sp. n.

- Fovea not oval, anterior margins diverging ( Figs 187View FIGURES 181–189, 196View FIGURES 190–197)................................................ typhon 

12(11) Areola elongated, anteriorly continuing into a chitinised crest ( Figs 160View FIGURES 154–161, 170View FIGURES 162–171)................................. signifer 

- Areola less elongated, anteriorly rounded ( Figs 7View FIGURES 1–9, 151View FIGURES 145–153)....................................................... 13

13(12) Epigyne as long as wide ( Figs 151View FIGURES 145–153, 165View FIGURES 162–171), lateral pockets nearly rounded ( Figs 152View FIGURES 145–153, 166View FIGURES 162–171)................... securifer  sp. n.

- Epigyne longer than wide, ( Figs 7View FIGURES 1–9, 22View FIGURES 18–29), lateral pockets more oval ( Figs 9View FIGURES 1–9, 23View FIGURES 18–29)............................... crassipes