Haplodrassus typhon ( Simon, 1878 )

Bosmans, Robert, Kherbouche-Abrous, Ourida, Benhalima, Souâd & Hervé, Christophe, 2018, The genus Haplodrassus Chamberlin, 1922 in the Mediterranean and the Maghreb in particular (Araneae: Gnaphosidae), Zootaxa 4451 (1), pp. 1-67: 49-50

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Haplodrassus typhon ( Simon, 1878 )

comb. n.

Haplodrassus typhon ( Simon, 1878)  , comb. n. stat. n.

Figs 181‒189View FIGURES 181–189, 194‒197View FIGURES 190–197, map 4

Drassus typhon Simon, 1878: 128  , pl. 16, fig. 20 (♂♀).

Drassodes macellinus: Simon, 1914: 209  (synonymy, here rejected).

Types : Female lectotype from France, without further locality, Coll. Simon ( MNHN AR9224View Materials); examined. 

Remarks. Drassus typhon  was described on the male and female sex by Simon in 1878. Later, he placed it in the synonymy of D. macellinus  , considering it “une variété non constante” ( Simon 1914). The only material of this species in the MNHN is a female labelled “AR9224 Drassodes typhon Simon E. Simon  det.”. It fits with the description of the epigyne given by Simon (1878) (“vide médian en ovale plus allongé”), which is one of the main differences to distinguish H. macellinus  from H. typhon  . We designate here this specimen as lectotype. The other examined material attributed by us to this species (see below) shows some variation but is treated here as the same species. The male and female described below were collected only once at the same site in Portugal by A. Grabolle and are considered conspecific. One further male was collected in the Bouches-du-Rhône department in France and several females without males in Algeria, Morocco, Spain and Italy.

Diagnosis. Males of Haplodrassus typhon  are easily distinguished by the elongated, pointed tibial apophysis ( Figs 186View FIGURES 181–189, 195View FIGURES 190–197) and the nearly straight embolic apophysis ( Figs 184View FIGURES 181–189, 194View FIGURES 190–197). Females are closely related to H. dentifer  sp. n., H. ovatus  sp. n. and H. rufipes  . They differ from H. ovatus  by the converging anterior margin of the fovea in H. ovatus  ( Fig. 105View FIGURES 98–107), diverging in H. typhon  ( Fig. 187View FIGURES 181–189), from H. dentifer  and H. rufipes  by the denticulate margin of the fovea ( Figs 40View FIGURES 30–40, 132View FIGURES 126–135).

Description. Measurements: Male (n=3): Total length 4.7‒6.0; carapace 2.18‒2.70 long, 2.0 wide. Female (n=9): Total length 6.2‒8.2; carapace 2.10‒2.90 long, 1.48‒2.30 wide.

Colour: Thoracic part of carapace yellowish brown, cephalic part reddish brown; legs yellowish brown; sternum reddish brown; abdomen pale brown, some specimens with traces of dorsal chevrons.

Male palp ( Figs 183‒186View FIGURES 181–189, 194‒195View FIGURES 190–197): Tibial apophysis elongated, strongly asymmetric, with pointed tip; embolic apophysis nearly straight, shorter than embolus, with rounded top.

Epigyne ( Figs 187‒189View FIGURES 181–189, 196‒197View FIGURES 190–197): Hood weekly sclerotized, separated from pro-fovea by a straight line; profovea much shorter than the elongated fovea, with anterior margins diverging; areola wide, with large lateral pockets.

Further material examined. AFRICA, ALGERIA, Annaba, Bône (= Annaba), 1♀, Bordj Bou Arreridj, 1♀, E. Simon leg., Coll  . Simon 6073 ( MNHN AR15715View Materials), El Bayadh, Le Kreider (N34°8’41” E0°4’9”), 1♀, E. Simon leg., CollGoogleMaps  . Simon 11933 ( MNHN AR15716View Materials), M'sila, Chott el Hodna, S. Baniou (N35°24'55” E4°20'39”), 400m, 2♀,GoogleMaps 

pitfalls in dunes in salt marsh, 1.VII.1988 ( CRB). TUNISIA, Medenine  , Djerba , 1♀, Coll. Simon 12475 ( MNHN AR15711View Materials)  . EUROPE, FRANCE, Bouches-du-Rhône, St-Antonin-sur-Bayon, Réserve Naturelle de Sainte Victoire (N43°31’44” E5°32’22”), 1♂, stones in low maquisGoogleMaps  , 4.V.2015 ( CJVK), Hérault, Brenas, La Lieude (N43°37’54” E3°16’10”), 1♀, stones in maquisGoogleMaps  , 13.V.2001, P. Oger leg. (CPO). SPAIN, Ciudad Real, Pozuelo de Calatrava (N38°53’6” W3°50’53”), 1♀, J. M. de la Fuente leg., CollGoogleMaps  . Simon 24173 ( MNHN AR15712View Materials), Granada, Sierra Elvira (N37°13'54” W 3°43'24”), 3♀, 17.VI.1908, E. Simon leg., CollGoogleMaps  . Simon 21738 ( MNHN AR15713View Materials), Murcia, Cartagena (N37°36’18” W0°59’10”), 3♀, E. Simon leg., CollGoogleMaps  . Simon 13414 ( MNHN AR15714View Materials). PORTUGAL, Algarve, Pereiro (N37°26’54” W7°35’46”), 2♂ 1♀, stones in dry, open grasslandGoogleMaps  , 21.IV.2012, A. Grabolle leg. ( CAG)  . ITALY, Sardinia, Oristano, Sinis (N39°53’5” E8°26’8”), 1♀, Marcialis leg. ( MCSNB)GoogleMaps  .

Distribution. N. Algeria, Tunisia, Spain, Portugal, S. France and Italy (map 4).


Museum National d'Histoire Naturelle


Universit� degli Studi di Cagliari














Haplodrassus typhon ( Simon, 1878 )

Bosmans, Robert, Kherbouche-Abrous, Ourida, Benhalima, Souâd & Hervé, Christophe 2018

Drassus typhon

Simon, 1878 : 128

Drassodes macellinus: Simon, 1914 : 209

Simon, 1914 : 209