Haplodrassus signifer ( C.L. Koch, 1839 )

Bosmans, Robert, Kherbouche-Abrous, Ourida, Benhalima, Souâd & Hervé, Christophe, 2018, The genus Haplodrassus Chamberlin, 1922 in the Mediterranean and the Maghreb in particular (Araneae: Gnaphosidae), Zootaxa 4451 (1), pp. 1-67: 42-44

publication ID

https://doi.org/10.11646/zootaxa.4451.1.1

publication LSID

lsid:zoobank.org:pub:5DCE18B3-9DBA-48F5-AEF7-90A50B984EA5

persistent identifier

http://treatment.plazi.org/id/DE5087A6-C03C-B31D-FF39-718EFEADFAC5

treatment provided by

Plazi

scientific name

Haplodrassus signifer ( C.L. Koch, 1839 )
status

 

Haplodrassus signifer ( C.L. Koch, 1839) 

Figs 154‒161View FIGURES 154–161, 167‒171View FIGURES 162–171

Drassus signifer C. L. Koch, 1839: 31  , fig. 452 (♀).

Drassodes troglodytes: Simon 1914: 122  , 140, 209, Figs 249‒250View FIGURES 241–251.

Haplodrassus signifer: Grimm 1985: 147  , figs 146‒148, 170‒171; Levy 2004: 19, figs 46‒52.

Types. Holotype female of Drassus signifer  from Bohemia; not examined. 

Diagnosis. Males of this species are easily recognized by the presence of a distinct crest on the sigmoid embolic apophysis of the palp ( Figs 158View FIGURES 154–161, 164View FIGURES 162–171) and the rounded dilatation of the tibial apophysis ( Figs 159View FIGURES 154–161, 168View FIGURES 162–171). Females resemble H. securifer  sp. n. but can be distinguished by the straight anterior margin of the pro-fovea ( Figs 151View FIGURES 145–153, 165View FIGURES 162–171), curved in H. signifer  ( Figs 160View FIGURES 154–161, 170View FIGURES 162–171) and the more rounded lateral pockets ( Figs 166View FIGURES 162–171 versus 171).

Description. Measurements: Male (n=20): Total length 4.7‒7.4; carapace 2.1‒3.4 long, 1.7‒2.7 wide. Female (n=20): Total length 5.1‒11.8; carapace 2.1‒5.5 long, 2.1‒5.5 wide.

Colour: Carapace reddish brown with darker cephalic part and radiating striae; legs yellowish brown; abdomen dark grey to black, often with obscure paler chevrons. Spinulation: General pattern, but ventral spines on Mt I often absent.

Male palp ( Figs 156‒159View FIGURES 154–161, 167‒169View FIGURES 162–171): Tibial apophysis distinctly constricted distally, then widening into a rounded knob ( Figs 159View FIGURES 154–161, 168View FIGURES 162–171); embolic apophysis S-shaped, distal part with a distinct crest ( Figs 158View FIGURES 154–161, 164View FIGURES 162–171).

Epigyne ( Figs 160‒161View FIGURES 154–161, 170‒171View FIGURES 162–171): As long as wide; hood strongly recurved laterally; pro-fovea short by the prolonged anterior margin of the areola, half the length of the fovea ( Figs 160View FIGURES 154–161, 170View FIGURES 162–171); fovea often with median groove; lateral pockets reniform ( Figs 161View FIGURES 154–161, 171View FIGURES 162–171).

Citations in the Maghreb. TUNISIA, Tunis, La Mohammedia ( Pavesi 1880), Tunis ( Pavesi 1884), Kairouan, Kairouan ( Pavesi 1880).

Further material examined. AFRICA, ALGERIA, Tébessa, Tébessa, 1300m, forêt de Bekkaria (N35°21'20” E8°14'40”), 2♂, pitfalls in Pinus halepensis  forest, 1.V.1989 ( CRB)GoogleMaps  . MOROCCO, Béni Mellal-Khénifra, Aguelmane Azigza, 1♂ 1♀, 6.IV.1950, Ch. Alluaud leg. ( ISRM)  ; Rabat-Salé-Kénitra, Oulmès S., ravin de l’Oued Assellalou (N33°25’25” W 6°00’ 10), 1♀, 11.III.1955, J. B. Panouse leg. ( ISRM)GoogleMaps  . EUROPE, SPAIN, Alava, Salvatierra (N42°51'5” W2°23'28”), 1♂ 2♀, stones in low maquis, 6.IV.1992 ( CRB)GoogleMaps  , Avila, Puerto del Pico (N40°19'16” W5°0'46”), 1300m, 1♀, 15.V.1992, P. Poot leg. ( CRB)GoogleMaps  , Puerto de Menga (N40°28'36” W5°0' 42""), 1570m, 1♀, stones in grassland, 1.IV.1997 ( CRB)GoogleMaps  , Burgos , Puerto de la Brugula (N42°25'49” W3°30'11”), 990m, 2♀, stones in Pinus  forest, 31.III.1997 ( CRB)GoogleMaps  , Cantabria, Puerto de San Glorio (N43°4'1” W4°45'51”), 1500m, 1♂ 2♀, pitfalls in wasteland, 17.VII.1985 ( CRB)GoogleMaps  , Ciudad Real, Pozuelo de Calatrava (N38°53’6” W3°50’53”), 2♀, J. M. de la Fuente leg, Coll. Simon 24235 ( MNHN AR15705View Materials), Gerona , Collada de Toses (N42°19'50” E2°2'10”), 1800m, 3♂, pitfalls in dry Pinus  forest, 10.VII.1991 ( CRB)GoogleMaps  , Málaga, Casares (N36°26'48” W5°17'8”), 2♀, V.1986, J. De Beir leg. (CRB), Palencia, Fuentes de Nava, Laguna de la Nava (N39°2’58” W3°41’47”), 1♂ 1♀, IV.‒XI.1996, U. Stengele leg. ( CRB)GoogleMaps  , Rioja, Navarrete (N42°25'41” W2°33'49”), 420m, 1♂ 1♀, stones in grassland, 7.IV.1992 ( CRB)GoogleMaps  , Segovia, 10 km E. Casla (N41°7'20” W3°41'52”), 1100m, 1♀, stones in grassland, 31.III.1997 ( CRB)GoogleMaps  , Toledo, Talavera la Nueva (N39°55'11” W4°52'22”), 370m, 1♂, litter in forest along Rio Tajo, 1.IV.1997 ( CRB)GoogleMaps  . FRANCE, mixed unknown localities (‘ Gallia Meridionalis’), one of them Alpes-Maritimes, Menton, 2♂ 1♀, as Drassodes macellinus hebes, Coll. Simon 2083  ( MNHN AR15708View Materials), Haute-Corse, Calacuccia, Lozzi, road to Monte Cinto (N42°20'46” E9°0'25”), 1200m, 2♀, stones in maquis, 25.V.1995 ( CRB)GoogleMaps  , 10 km from Calvi (N42°33’57” E8°45’25”), 1♂ 2♀, pitfalls in maquis, 15. V.1982 ( CRB)GoogleMaps  , Col de Vergio (N42°17'24” E8°52'39”), 1460m, 3♂ 1♀, stones in Juniperus  forest, 24.V.1995 ( CRB)GoogleMaps  , Gorges de Restonica (N42°14'28” E9°10'52”), 1300m, 1♂, stones in Pinus laricio  forest, 26.V.1995 ( CRB)GoogleMaps  , E. Urtaca (N42°35'40” E9°9' 57'), 1♂, stones in garrigue, 23.V.1995 ( CRB)GoogleMaps  , Haute -Savoie, Saint Julien-Mont-Denis (N45°14'48” E6°25'30”), 750m, 1♀, pitfalls in grassland, 6.VI.1986 (CRB), Pyrénées-Orientales, Mont Canigou, Escala de l’Ours (N42°32'58” E2°29'40”), 1400m, 1♂, 5.VII.1991 ( CRB)GoogleMaps  , Col de Puymorens , 1850m, 2♂ 1♀, pitfalls in dry grassland, 11. VI.1991 ( CRB)  . ITALY, Sardinia, Carbonia-Iglesias, Fluminimaggiore S. (N39°24'38” E8°29'26”), at entrance of Grotta su Manua , 200m, 1♂, seeving Quercus  litter, 15.IV.2014 ( CRB)GoogleMaps  , Nuoro , Orgosolo N. (N40°13'6” E9°22'47”), 610m, 1♀, seeving Quercus  litter, 18.IV.2014 ( CRB)GoogleMaps  , Sassari , Alghero W., Parco di Porto (N40°36'59” E8°9'51”), 1♂ 1♀, 10.IV.2014 ( CRB)GoogleMaps  . GREECE, Attiki-Saronic Islands, Attiki, Marathon, Enoee ruins (N38°9'28” E23°56'1”), 80m, 1♂ 1♀, stones in phrygana, 19.IV.2000 ( CRB)GoogleMaps  , Oros Imittos, W. Peania (N37°56'47” E23°50'0”), 450m, 1♀, 20.IV.2000 ( CRB)GoogleMaps  , Crete, Sitia , 2♀, as H. severus, Grimm  det. ( MCSNB)  , Ipeiros, Ioannina, Vikos-Aoos national park, Astraka (N39°57'54” E20°47'31”), 1910m, 2♀, pitfalls in grassland, 12.VII.2007, B. Vandenberghe leg. ( CRB)GoogleMaps  , Evvoia- Voroies Sporades, Evvoia, Papades, Aghia Anna (N38°55'44” E23°21'51”), 550m, 1♀, stones in grassland, 12.V.2001 (CRB), Evvoia, Prokopi-Nea Pagontas road (N38°42'38” E23°30'57”), 700m, 1♂, stones in grassland, 10.V.2001 (CRB), Kyklades: Iraklia (N36°50’15” E25°27’21”), 50m, 1♀, on walls of house, 6.III.2016, 1♀, 9.IV.2017, 1♂, in pitfals in phrygana, 20.IV.2017, Y. Gavalas leg. (CYG)GoogleMaps  ; Makedonia, Halkidiki, Monopigado (N40°26'4” E23°8'5”), 400m, 1♂, pitfalls in Quercus  forest, 22.III ‒12.VI.2004, L. Provoost leg. ( CRB)GoogleMaps  , Thessaloniki, NE. Thessaloniki (N40°42'20” E22°58'58”), 200m, 1♂, pitfalls in dry Pinus  forest, 7.II ‒6.VI.2004, L. Provoost leg. ( CRB)GoogleMaps  , Peloponnisos, Argolis, Achladokambos (N37°32'44” E22°31'10”) 1000m, 2♀, under stones, 26.IV.2016 ( CRB)GoogleMaps  , Korinthia, Lake Stymphalia , (N37°50'39” E22°27'15”), 615m, 1♂, pitfalls under Quercus  bushes on slope to lake, 20.IV ‒27.IV.2016 ( CRB)GoogleMaps  , Sterea Elada, Etolio-Akarnania, Paleros S., Agios Dimitriou (N38°46'6” E20°53'6”), 50m, 2♀, stones in quarry, 17.IV.2000 ( CRB)GoogleMaps  , Stanos S., lake Amvrakia (N38°46'51” E21°11'17”), 20m, 1♂, stones in olive groove, 15.IV.2000 ( CRB)GoogleMaps  , Fokida, Lake Mornou (N38°31'06” E22°09'06”), 440m, 1♀, 25.IV.2015GoogleMaps  , P. Ponel leg. (CPP), Voiotia, Oros Parnassos, Kalivia Arachovas (N38°31'24” E22°32'49”), 900m, 1♀, pitfalls in grassland, 18.IV.2000 (CRB), Thiva (N38°19’30” E23°19’08”), 200m, 1♀, 1998, L. Picard leg. ( CRB)GoogleMaps  . CYPRUS, Famagusta, Büyükkonuk ( Konik ) (N35°24'32” E33°59'28”), 150m, 1♂, pitfalls in Pinus  forest, III ‒V.2011 ( CRB)GoogleMaps  , Yenibogaziçi ( Agios Sergios ) (N35°12'3” E35°53'43”), 30m, 1♂, stones in phrygiana, 17.III.2011, 1♂, in pitfalls at same site, 15‒ 27.V.2011 ( CRB)GoogleMaps  , Kyrenia, Kalkani ( Kalo Chorio ) (N35°15'27” E33°2'22”), 100m, 1♀, stones in grassland, 20.III.2011 ( CRB)GoogleMaps  , Vouni ruins (N35°10'11” E32°46'24”), 250m, 1♀, under stones, 20.IIII.2011 ( CRB)GoogleMaps  , Pafos, Kouklia-Pissoura road (34°40'27” E32°37'06”), 150m, 1♀, stony slope to the sea, 22.III.2011 ( CRB)  , Nicosia, Prodromos E., near Prodromos dam (N34°56'58” E32°50'39”), 1550m, 1♀, stones in Pinus  forest, 23.V.2001 ( CRB)GoogleMaps  .

Distribution. Holarctic. In the Nearctic it is known from southern Canada, to central-S Mexico, in the Palaearctic up to 70°. In the Maghreb, the species appears to be rare. It is known from the NE of Algeria, Morocco, Tunisia and Libya.

MNHN

Museum National d'Histoire Naturelle

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Gnaphosidae

Genus

Haplodrassus

Loc

Haplodrassus signifer ( C.L. Koch, 1839 )

Bosmans, Robert, Kherbouche-Abrous, Ourida, Benhalima, Souâd & Hervé, Christophe 2018
2018
Loc

Drassus signifer

C. L. Koch, 1839 : 31
Loc

Drassodes troglodytes:

Simon 1914 : 122
Loc

Haplodrassus signifer: Grimm 1985 : 147

Grimm 1985 : 147
Levy 2004 : 19