Proagoderus mabuensis Daniel, Josso, Nganhane & Strümpher, 2024

Daniel, Gimo M., Josso, Jean-François, Nganhane, Isildo De N. & Strümpher, Werner P., 2024, Dung beetle fauna from Mount Mabu, Mozambique. Part 3: A new species of Proagoderus van Lansberge, 1883 with an annotated checklist of the Proagoderus dives (Harold, 1877) species-group (Coleoptera: Scarabaeidae: Scarabaeinae), Zootaxa 5406 (1), pp. 175-189 : 176

Proagoderus mabuensis Daniel, Josso, Nganhane & Strümpher , new species urn:lsid:zoobank.org:act:1CF19F0B-36D4-4D23-93B1-6F8C6855F46A

( Figs. 1–5 )

Type locality. Transect 4 (16°16’38”S, 36°22’45”E), Mount Mabu , Zambézia Province, Mozambique ( Fig. 10 ) .

Type material. Holotype (♂ TMSA): “MOZ [ MOZAMBIQUE]: Zambézia [Province] | Mabu Forest, Mt. Mabu | Trans. [ Transect ] 4 [on foothills] | 16°16’38”S, 36°22’45”E | 15–19.iv. 2022, 1350 m | Daniel, Nganhane & Strumpher || Hum. [Human] dung pitfall trap | Afromontane | Forest || HOLOTYPE Proagoderus | mabuensis | Daniel, Josso, | Nganhane, Strümpher, 2023 [red cardstock]” ( Fig. 3 ) . Paratypes. (28 TMSA) : 15♂♂, 13♀♀: same data, except: “PARATYPE Proagoderus | mabuensis Daniel, Josso | Nganhane, Strümpher, 2023”; (283 BMSA): (3♂♂, 5♀♀ BMSA (C) 140152–140152, 140297–140335): “MOZ [ MOZAMBIQUE]: Zambézia [Province] | Mabu Forest, Mt. | Mabu, Trans. [Transect] 1 | 16°16’37’’S, 36°22’49”E | 15–19.iv. 2022, 1355 m || Hum. [Human] dung pitfall trap |Afromontane | Forest || Terres. [Terrestrial] Invert. [Invertebrates] Dept. [Department] | National Museum | P.O. BOX 266 | Bloemfontein 9300 | South Africa [blue cardstock] || PARATYPE | Proagoderus mabuensis | Daniel, Josso, | Nganhane & Strümpher, 2023 [yellow cardstock]”; (12♂♂, 10♀♀ BMSA (C) 140184–140190, 140271, 140279–140292): same data, except: “Mabu, Trans. [Transect] 2 | 16°16’38’’S, 36°22’51”E | 15–19.iv. 2022, 1334 m”; (6♂♂, 12♀♀ BMSA (C) 1400093–140109, 140182–140182): same data, except: “Mabu, Trans. [Transect] 3 | 16°16’37”S, 36°22’46”E | 15–19.iv. 2022, 1334 m”; (18♂♂, 25♀♀ BMSA (C) 140154–140173, 140193–140196, 140314–140317, 140338–140353): same data, except: “Mabu, Trans. [Transect] 4 | 16°16’38”S, 36°22’45”E | 15–19.iv. 2022, 1350 m”; (22 ♂♂, 46♀♀ BMSA (C) 140089–140091, 140118, 140120–140151, 140278, 140302– 140313, 140354–140375): same data, except: “Mabu, Trans. [Transect] 5 | 16°16’33”S, 36°22’44”E | 16–19.iv. 2022, 1360 m”; (20♂♂, 27♀♀ BMSA (C) 140110, 140112–140115, 140194–140235): same data, except: “Mabu, Trans. [Transect] 6 | 16°16’28’’S, 36°22’37”E | 16–19.iv. 2022, 1330 m”; (11♂♂, 20♀♀ BMSA (C) 140172–140181, 140254–140266): same data, except: “Mabu, Trans. [Transect] 7 | 16°16’51”S, 36°22’57”E | 16–19.iv.2022, 1400 m || Hum. [Human] dung pitfall trap | Dense woodland & | bamboo forest”; (25♂♂, 15♀♀ BMSA (C) 140236–140253, 1402318–140334, 1402376–140381): same data, except: “Mabu, Trans. [Transect] 10 | 16°16’45”S, 36°22’04”E | 17–20.iv.2022, 1200 m || Hum. [Human] dung pitfall trap | Afromontane | Forest”; (2♂♂, 4♀♀ BMSA (C) 140087, 140116–140117, 140191–140193): same data, except: “| Mabu Forest, Mt. | Mabu Foothills, Trans. [Transect] 12 | 16°17’04’’S, 36°23’43”E | 18–20.iv. 2022, 109 m || Hum. [Human] dung pitfall trap | Dense Woodland & Bamboo | Forest”; (4 FMNH): (1 ♂): “MOZ [ MOZAMBIQUE]: Zambézia [Province] | Mabu Forest, Mt. Mabu, Trans. [Transect] 3 | 16°16’37’’S, 36°22’46”E | 15–19.iv. 2022, 1334 m | Daniel, Nganhane & Strumpher || Hum. [Human] dung pitfall trap | Afromontane | Forest || Proagoderus van Lansberge 1883 | new species, det. GM | Daniel 2023 || PARATYPE | Proagoderus mabuensis | Daniel, Josso | Nganhane & Strümpher 2023 [yellow cardstock]”; (1♀): same data, except: “Mabu, Trans. [Transect] 4 | 16°16’38”S, 36°22’45”E | 15–19.iv. 2022, 1350 m”; (1♀): same data, except: “Mabu, Trans. [Transect] 4 | 16°16’38”S, 36°22’45”E | 15–19.iv. 2022, 1350 m”; (1♂) same data, except: “Mabu Forest, Mt. | Mabu Foothills, Trans. [Transect] 12 | 16°17’04”S, 36°23’43”E | 18–20.iv. 2022, 109 m || Hum. [Human] dung pitfall trap | Dense Woodland & Bamboo | Forest”; (6 JFJC): (1♂, 2♀♀): “MOZ [ MOZAMBIQUE]: Zambézia [Province] | Mabu Forest, Mt. Mabu, Trans. [Transect] 3 | 16°16’37”S, 36°22’46”E | 15–19.iv. 2022, 1334 m | Daniel, Nganhane & Strumpher || Hum. [Human] dung pitfall trap | Afromontane | Forest || PARATYPE | Proagoderus mabuensis | Daniel, Josso | Nganhane & Strümpher 2023 [yellow cardstock]”. (3♀♀): same data, except: | Mabu Forest, Mt. Mabu, Trans [Transect]. 6 | 16°16’28”S, 36°22’37”E | 15–19.iv. 2022, 1330 m ”.

Figures

FIGURE 10. Distribution of Proagoderus mabuensis Daniel, Josso, Nganhane & Strümpher, new species. Image reproduced from Daniel et al. (2023b).

FIGURES 1–5. Proagoderus mabuensis Daniel, Josso, Nganhane & Strümpher, new species. 1, habitus (holotype, male, TMSA), dorsal view; 2, forebody of a female paratype (TMSA); 6, holotype labels; 4–5, aedeagus (holotype) in lateral and dorsal views.