Cladiscus weyersi Kraatz, 1899

Murakami, Hiroyuki, 2020, Review of the genus Cladiscus (Coleoptera: Cleridae) from Japan and Taiwan, with descriptions of two new species, Acta Entomologica Musei Nationalis Pragae 60 (2), pp. 475-492 : 480-481

Cladiscus weyersi Kraatz, 1899

( Figs 2 , 16 , 28, 35, 52–58, 126)

Cladiscus weyersi Kraatz, 1899: 213 [original description].LOΗDΕ (1900): 17 [listed]; S CΗΕΝΚLΙΝG (1910): 10 [listed]; CORΡORΑΑL +| VΑΝ DΕR WΙΕL (1949): 196 [key to species]; CORΡORΑΑL (1950): 24 [listed].

Type locality. Indonesia, Sumatra (after KRΑΑΤΖ 1899).

Specimens examined. TAIWAN: PIΝGτUΝG COUΝτ Y: Shouka, Shihtzu Township, 11.v.2014, 2 ♁♁, Jiafong Chen leg. ( JCCT).

Differential diagnosis. This species is similar to C. mindanensis Chapin, 1924 from the Philippines, but it is distinguished by the following characters (CORΡORΑΑL +| VΑΝ DΕR WΙΕL 1949): scutellum red; head and pronotum finely and sparsely punctured.

Supplementary description. Male ( Fig. 2 ). Head, pronotum, and antennomeres I and II brownish red; antennomeres III to XI, elytra, and legs brownish black.

Antennae ( Fig. 16 ) longer than total length of head and pronotum combined; antennomere I swollen and bent; II compact; III to X pectinate, with branches arising at base of antennomeres; XI fusiform.

Pronotum smooth at apex; postcoxal projections ( Fig. 35 ) long; prosternal process not extending contact point of postcoxal projections. Elytra with rows of large, deep, asetigerous punctures in basal half; apical margins finely indented. Mesoventrite with sparse setigerous punctures except for central portion. Metaventrite with evenly arranged fine setigerous punctures except for central portion; metaventral anterior process elongate, as wide as mesoventral posterior process. Metendosternite ( Fig. 52 ) without furcal laminae; process extending anteriorly, tapered; furcal arms almost straight, extending anterolaterally.

Protibiae ( Fig. 28 ) gently widening from basal 1/3 to apex, weakly curved; meso- and metatibiae almost straight. Tibial spur formula 0–0–0.

Apical margin of abdominal ventrite V broadly and weakly emarginate. Apical margin of pygidium ( Fig. 53 ) straight; struts short. Apical margin of ventrite VI ( Fig. 54 ) slightly emarginated. Spicular fork ( Fig. 55 ) with intraspicular plate, which is 1/2 as long as spicular fork.

CM ( Figs 56–58 ) covered with large spines and fine spines, 1/4 as long as tegmen. Tegmen shorter than phallus; phallobasic apodeme fused with phallobasic struts. Phallus widest at 1/3, thence tapered apically.

Female. Unknown.

Measurements and ratios. Male (n Y 2): BL: 8.08–8.82 mm; PL: 2.08–2.28 mm; PWA: 1.40 mm; PWM: 0.73–0.75 mm; PWB: 0.93–0.98 mm; EL: 6.00– 6.55 mm; EW: 1.50–1.63 mm; EyD: 0.75–0.78 mm; EyW: 0.38–0.40 mm; EL/PL: 2.88–2.89; EW/PWA: 1.07–1.16; EW/PWM: 2.07–2.17; EW/PWB: 1.62–1.67; PWA/PWB: 1.44–1.51; PWA/PWM: 1.87–1.93; PWB/PWM: 1.28–1.30; EyD/ EyW: 1.94–2.00.

Distribution ( Fig. 126 ). Taiwan (new record); India (CORΡORΑΑL +| VΑΝ DΕR WΙΕL 1949); Indonesia: Sumatra (KRΑΑΤΖ 1899), Java (CORΡORΑΑL +| VΑΝ DΕR WΙΕL 1949).

Figures

Fig 126. Distribution map of the species of Cladiscus from Japan and Taiwan.

Figs 1–13. Habitus and head of Cladiscus spp. (1–5, 6, 8, 10, 12, male; 7, 9, 11, 13, female). 1 – Cladiscus obeliscus Lewis, 1892; 2 – C. weyersi Kraatz, 1899; 3 – C. sauteri Schenkling, 1912; 4 – C. terminalis Schenkling, 1912; 5 – C. pallidicornis Corporaal & van der Wiel, 1949; 6, 7, 10, 11 – C. hachijoensis sp. nov. (6, 10, holotype; 7, 11, paratype); 8, 9, 12, 13 – C. liaoi sp. nov. (8, 12, holotype; 9, 13, paratype). Scale bars: 2.0 mm.

Figs 14–26. Antennae (14, 16, 17, 19, 21, 23, 25 – male; 15, 18, 20, 22, 24, 26 – female). 14, 15 – Cladiscus obeliscus Lewis, 1892; 16 – C. weyersi Kraatz, 1899; 17, 18 – C. sauteri Schenkling, 1912; 19, 20 – C. terminalis Schenkling, 1912; 21, 22 – C. pallidicornis Corporaal & van der Wiel, 1949; 23, 24 – C. hachijoensis sp. nov.; 25, 26 – C. liaoi sp. nov.

Figs 27–39. Male protibiae (27–33) and prothorax in ventral view (34–39). 27, 34 – Cladiscus obeliscus Lewis, 1892; 28, 35 – C. weyersi Kraatz, 1899; 29, 36 – C. sauteri Schenkling, 1912; 30, 37 – C. terminalis Schenkling, 1912; 31, 38 – C. pallidicornis Corporaal & van der Wiel, 1949; 32, 39 – C. hachijoensis sp. nov.; 33 – C. liaoi sp. nov. Scale bars: 0.5 mm.

Figs 52–58. Metendosternite, terminal parts and genitalia in male of Cladiscus weyersi Kraatz, 1899. 52 – metendosternite; 53 – pygidium; 54 – ventrite VI; 55 – spicular fork; 56–58 – male genitalia in lateral (56), ventral (57), and dorsal (58) views.