Ins pleuralis (Williston) Evenhuis, 2020

Evenhuis, Neal L., 2020, A new genus for Painter & Painter’s Villa “ celer ” - group in the New World (Diptera: Bombyliidae), Zootaxa 4748 (2), pp. 296-314 : 309

Ins pleuralis (Williston) , new combination

( Figs. 2 , 18 , 28 )

Anthrax pleuralis Williston, 1901: 282 .

Villa (Hemipenthes) pleuralis (Williston) . Painter & Painter 1962: 121.

Hemipenthes pleuralis (Williston) . Hull 1973: 373.

Types. Holotype ♀ from: MEXICO: Guerrero: Vente de Zopilote , 2800 ft., H.H. Smith in BMNH. Type examined by Painter & Painter (1962: 121).

Diagnosis. Similar to I. leucocephala (Wulp) in wing venation with the anal cell not completely infuscated, but can be separated from it by the infuscation fading posteriorly (dimidiate infuscation in I. leucocephala ), and the infuscation in cell r2+3 extending past origin of R 2+3 almost to the origin of R 4 ( Fig. 18 ) (this infuscation not extending much past origin of R 2+ 3 in I. leucocephala ).

Material examined. MEXICO: Colima: 1♂, 1♀, Colima, 1500 ft., 25Aug 1962, R . H. & E.M. Painter ( USNM) ; Guerrero: 1♂, 10 mi. S. Taxco, 14 Aug 1962, N. Marston–21 ( USNM) ; Morelos: 1♂, 5 m S. Cuernavaca, 16 Aug 1962, N. Marston–30. ( USNM) .

Remarks. Ávalos-Hernández (2009) indicated that there were similarities in external features of Ins pleuralis and I. galathea (both treated there as Hemipenthes species) but would require further study of genitalia to verify ( I. galathea was not within the scope of study of Hemipenthes by Ávalos-Hernández). The wing infuscation alone suffices to separate the two as I. pleuralis is has the infuscation extending almost to the wing tip ( Fig. 18 ). However, a comparison of the male genitalia shows that the ventral blade-like process of the parameral sheath is small and subtriangular in lateral view in I. pleuralis ( Fig. 28 ), while that in I. galathea is longer in lateral view, extending from tip of parameral sheath to level of apex of distiphallus ( Fig. 24 ); in the latter respect, I. galathea appears closer to I. martinorum ( Fig. 26 ).

Distribution. Mexico (Chiapas, Colima, Guerrero, Morelos, Nayarit, Puebla, Sinaloa, Tamaulipas).

Figures

FIGURES 1–2. Habitus views. 1, Hemipenthes sinuosa (Wiedemann) dorsal view; 2, Ins pleuralis, dorsal view.

FIGURES 10–19. Hemipenthes wing. 10, H. sinuosa. 11–17, Ins wings. 11, I. celeris; 12, I. curta; 13, I. constituta; 14, I. ignea; 15, I. martinorum; 16, I. minas; 17, I. pectorcolumbo, n. sp.; 18, I. pleuralis; 19, I. zanouts, n. sp.

FIGURES 24–28. Ins, male genitalia. 24, I. constituta, A, distiphallus and parameral sheath, lateral view, B. parameral sheath dorsal view. 25, I. ignea, A, distiphallus and parameral sheath, lateral view, B. parameral sheath dorsal view. 26, I. martinorum, A, distiphallus and parameral sheath, lateral view, B. parameral sheath dorsal view. 27. I. minas, A, distiphallus and parameral sheath, lateral view, B. parameral sheath dorsal view. 28 I. pleuralis, A, distiphallus and parameral sheath, lateral view, B. parameral sheath dorsal view.