Satsuma albida (H. Adams, 1870)

Wu, Shu-Ping, Hwang, Chung-Chi & Lin, Yao-Sung, 2008, Systematic revision of the arboreal snail Satsuma albida species complex (Mollusca: Camaenidae) with descriptions of 14 new species from Taiwan, Zoological Journal of the Linnean Society 154 (3), pp. 437-493 : 442

SATSUMA ALBIDA SPECIES COMPLEX

Diagnosis: The colour of shell, periostracum and soft body is white to yellowish, never brownish as seen in other Satsuma . The shell is conical with shell height/ shell diameter ratios of 1.0–1.8.

Description: Shell dextral, conical, thin to thick, small to medium size (see Table 2 for shell measurements). Shell white or yellowish with or without coloured bands and stains; shell of some species with greenish or golden tint. Bands highly polymorphic, pink, reddish purple or dark purple, variable in width and shape ( Fig. 2 ). Whorls not obviously expanded. Periphery bluntly angulated to keeled. Base expanded or flattened. Surface glossy with curved striae and thin periostracum. Aperture diagonally tetragonal or ovate-lunate. Peristome thin, reflected. Pre-apertural constriction behind outer and inferior lips present on some species. Insertion of columellar lip dilated, not connected with insertion of outer lip. Callus between insertions thin, present only on some species. Umbilicus narrow, half to mostly covered by dilated columellar lip. Colour of soft body in accordance with shell, either white or yellowish. Longitudinal pedal stripe of grey colour present on ventral side of foot in white-coloured individuals. Yellow-shelled species do not exhibit such pedal stripe.

The vagina and penis can be divided into three segments, namely proximal, middle and distal parts for convenience of description. Boundaries between parts can be recognized by apparent changes of the external and internal morphologies, such as the strength of parts and the number, shape and strength of internal folds. However, some species showed no apparent boundary between these parts. The verge around the epiphallic pore forms two to three strong pilasters (called caecal pilasters hereafter in this article) tapering to the apex of the penial caecum and usually fused as a single principal pilaster extending distally to the proximal penis.

Bursa stalk long with 8–15 smooth and straight internal folds. Free oviduct short. Proximal vagina muscular, swollen, smooth with strong internal folds. Middle vagina constrictive with wiggly and dense internal folds in some species. Distal vagina and atrium villous, dull externally with fine corrugations internally. Flagellum slender or conical. Penis and epiphallus bundled by thin and semi-transparent penial muscle. Proximal penis absent, short or long, with variable morphology. Middle and distal penis thin, slender with strong pilasters to fine wrinkles internally. In some species, a few to numerous variably sized, tiny, flexible and nearly regular tetrahedral spinules were born on the surfaces of pilasters in part of the penial caecum, penis and vagina. A representative figure of the spinules is supplied only for Satsuma mollicula as the morphology of spinules is similar between species under the stereo microscope.

Figures

Figure 2. Band formula and colour stain demonstrating polymorphism of banding patterns. Numbers 1–8 show the position of the band. A ‘0’ means no bands or stains present. Digits in parentheses denoted a smear displayed between and connected with the bands or stains. See text for definition of bands and stains in detail.