Novalena intermedia (Chamberlin & Gertsch 1930),

Maya-Morales, Julieta & Jiménez, María Luisa, 2017, Revision of the funnel-web spider genus Novalena (Araneae: Agelenidae), Zootaxa 4262 (1), pp. 1-88: 20-24

publication ID

http://doi.org/ 10.5281/zenodo.571683

publication LSID

lsid:zoobank.org:pub:894795FB-ED38-4132-92B9-6AC8C3E41566

persistent identifier

http://treatment.plazi.org/id/E45787C5-FF92-FFEC-E1A1-6190FE05FC69

treatment provided by

Plazi

scientific name

Novalena intermedia (Chamberlin & Gertsch 1930)
status

 

Novalena intermedia (Chamberlin & Gertsch 1930) 

Figures 8–11View FIGURE 8View FIGURE 9View FIGURE 10View FIGURE 11; Map 1

Agelena intermedia Chamberlin & Gertsch 1930: 143  , figure 28 (♂) Agelena (Hololena) intermedia Exline 1938: 28  , plate 4, figures 35–36 (♂ ♀) Novalena intermedia Chamberlin & Ivie 1941: 9  (transferred from Hololena  to generic nomen nudum) Novalena intermedia Chamberlin & Ivie 1942: 225  , figures 54–55 (♂ ♀) Novalena pina Chamberlin & Ivie 1942: 226  , figure 56 (♀) NEW SYNONYMY Novalena intermedia Lehtinen 1967: 446  , figures 223, 225 (♂ ♀) Novalena intermedia Roth & Brame 1972: 46  , figure 65 (♂)

Novalena pima Roth & Brame 1972: 46  , figure 64 (♀) (incorrect spelling) Novalena intermedia Roth 1982: 7  –9, figure 65 (♂)

Novalena pima Roth 1982: 7  –9, figure 64 (♀)

Novalena intermedia Roth 1985  : B1–8, figure 65 (♂)

Novalena pima Roth 1985  : B1–8, figure 64 (♀)

Novalena intermedia Roth 1994: 54  , figure 65 (♂)

Novalena pima Roth 1994: 54  , figure 64 (♀)

Types. Holotype male of N. intermedia  : USA: California: La Honda , 37°20′N, 122°17′W, 4.XI.1922, J.C. Chamberlin, deposited at AMNH (examined)GoogleMaps  . Holotype female of N. pina  : USA: California: Echo Lake , 38°08′N, 120°52′W, VII.1934, L.W. Saylor, deposited at AMNH (examined).GoogleMaps 

Synonymy justification. The overall appearance of the epigynum of N. pina  is as in N. intermedia  , in shape and position of spurs, copulatory ducts, primary and secondary spermathecae, as well as the protrusion of anterior margin of atrium.

Diagnosis. The males of N. intermedia  distinguish from all species in having the RTA with dorsal projection large and cupped ( Figs. 10View FIGURE 10 c, 11b). The females are similar to N. ajusco  , N. alvarezi  , N. dentata  , and N. leonensis  , but differ from those species in having the atrium with anterior margin not sclerotized ( Fig. 11View FIGURE 11 c).

Description. Male (holotype): Carapace light brown. Chelicerae dark brown, condyles brown. Endites brown with white tips. Labium dark brown with white tip. Sternum brown. Legs with femur and tarsus light brown, patella and metatarsus brown. Three weak rings on femur. Opisthosoma yellow with several brown spots, reddish foliage, and lateral yellow spots. Spinnerets light brown. Total length 9. Carapace length 4.6, width 3, cephalic region width 1.5, ocular region width 0.9. Chelicerae with two to three retromarginal teeth; basal segment length 1.5, fang length 0.8. Labium slightly wider then long (0.5/0.4). Endites slightly convergent (distance at their base compared at their tips 0.5/0.3). Sternum longer than wide (2/1.8). ALS separated by more their basal diameter (0.4/0.3), PLS with distal segment shorter than basal segment (0.3/0.5). Leg lengths: I—femur 3.5/ patella-tibia 4.65/ metatarsus 3.5/ tarsus 2.4; II—3.65/ 4.6/ 3.5/ 2; III—3.5/ 4.2/ 3.8/ 2; IV—4.2/ 5.4/ 5.1/ 2.4. Spination: femur dorsal I—1-2 -3/ II— 1-2 -3/ III—1-2 -3/ IV—1-2 -3; patella I—dorsal 1-1/ prolateral 1/ retrolateral 0; II—1-1 / 1/ 0; III—1-1 / 1/ 1; IV—1- 1 / 1/ 1; tibia I—dorsal 0/ ventral 2-2-2/ prolateral 1-1-0/ retrolateral 0; II—0/ 2-2-2/ 1-1-0/ 0; III—1-1 -0/ 2-2-2/ 1- 1-0/ 1-1-0; IV—1-1 -0/ 1-1-1-2/ 1-1-0/ 1-1-0; metatarsus I—0/ 2-2-2/ 0-1-1/ 0-1-1; II—1 -0-1/ 2-2-2/ 0-1-0/ 0-1-0; III—2-1 -2/ 2-2-2/ 0-1-1/ 0-1-0; IV—3-2 -2/ 2-1-1-2/ 0-1-1/ 0-0-1. Number of trichobothria on tarsus: I—7, II—6, III—7, IV—6. Number of dorsal spines on pedipalp: femur 3, tibia 3. Cymbium length 1.3, width 0.88. Embolus curved; conductor with ventral projection distally notched; tegular median process as a pointed projection; dorsal projection of RTA heavy and cupped ventrally; ventral projection short ( Figs. 10View FIGURE 10 a, c, e, 11a, b).

Female (allotype) of N. intermedia  : Carapace yellow with two longitudinal lines behind PME. Chelicerae dark brown, condyles light brown. Endites and labium light brown with white tips. Sternum light brown. Legs light brown, metatarsus dark brown. Three rings on femur, one on tibia III. Opisthosoma similar to male holotype. Spinnerets light brown. Total length 9. Carapace length 4.4, width 3, cephalic region width 1.6, ocular region width 1.1. Chelicerae with three retromarginal teeth; basal segment length 1.7, fang length 0.8. Labium longer than wide (0.6/0.5). Endites slightly convergent (distance at their base compared at their tips 0.5/0.4). Sternum longer than wide (2.2/1.8). Opisthosoma longer than wide (5.3/3). ALS separated by less their basal diameter (0.25/0.3), PLS with distal segment as long as basal segment (0.5/0.5). Leg lengths: I—femur 3.2/ patella-tibia 4.25/ metatarsus 2.9/ tarsus 1.9; II—3.1/ 3.9/ 2.7/ 1.7; III—3 / 3.6/ 3.1/ 1.5; IV—3.8/ 4.7/ 4.2/ 2. Spination: femur dorsal I—1-2 -3/ II—1-3 -3/ III—1-3 -3/ IV—1-2 -3; patella I—dorsal 1-1/ prolateral 1/ retrolateral 0; II—1-1 / 1/ 0; III—1-1 / 1/ 1; IV—1-1 / 1/ 1; tibia I—dorsal 0/ ventral 2-2-2/ prolateral 1-1-0/ retrolateral 1-1-0; II—0/ 2-2-2/ 1-1-0/ 1-1-0; III— 1-1 -0/ 1-2-2/ 1-1-0/ 1-1-0; IV—1-1 -0/ 1-1-0/ 1-1-0/ 1-1-2; metatarsus I—0/ 2-2-2/ 1-1-0/ 0; II—0/ 2-2-2/ 0-1-1/ 0- 1-1; III—3-2 -2/ 2-2-2/ 0-1-0/ 0-1-1; IV—3-2 -2/ 2-2-2/ 0-1-1/ 0-0-1. Number of trichobothria on tarsus: I—7, II— 7, III—7, IV—6. Dorsal spines on pedipalp femur: 3. Epigynal plate wider than long (1.9/0.7); atrium with anterior margin more protruded than posterior margin and lateral margins engrossed; anterior hyaline spurs separated by slightly more their basal width ( Fig. 11View FIGURE 11 c, e). Internal genitalia: copulatory ducts straight, primary spermathecae separated by less their width ( Figs. 10View FIGURE 10 b, d, f, 11d).

Variation. Males (n=10): body length 6–9; carapace length 3.5–4.6; patella-tibia I length 3.75–4.65. Females (n=10): body length 6.75–10.75; carapace length 3.38–4.75; patella-tibia I length 3.38–4.63. Other specimens with two to three retromarginal teeth on chelicerae.

Additional material examined. Allotype of N. intermedia  : USA: same data as holotype, 1 ♀ ( AMNH).GoogleMaps 

Other material: USA: Arizona: Pima  Co., Santa Rita Mountains, Madera Canyon , Round Camp , 23.III.1960, W.J. Gertsch, Ivie, and Schrammel, 1 ♂ ( AMNH)  . California: Alpine Co., Alpine , 10.IX.1959, W.J. Gertsch and V. Roth, 1 ♀ ( AMNH)  ; Calaveras Co., 8 km W Dorrington, 5.VIII.1953, W.J. Gertsch and J.W. Gertsch, 1 ♂ ( AMNH)  ; El Dorado Co., Echo Summit , 6.4 km SW Meyers, 2250 m, 9.IX.1959, W.J. Gertsch and V. Roth, 1 ♀ ( AMNH)  ; Fresno Co., Kings Canyon National Park, Cedar Grove , 13.IX.1959, V. Roth and W.J. Gertsch, 1 ♀ ( AMNH)  ; Fresno Co., Cherry Gap , 1829 m, 6.VIII.1955, R.X. Schick, 1 ♀ ( AMNH)  ; Mariposa Co., 9.7 km S Mather, 4.IX.1958, V. Roth, 1 ♀ ( AMNH)  ; Nevada Co., no date and collector data, 1 ♂ ( AMNH)  ; San Mateo Co., Woodside , 5.VII.1958, V. Roth and W.J. Gertsch, 2 ♀ ( AMNH)  ; Shasta Co., Hatchet Creek , 1.VIII.1953, W.J. Gertsch and J.W. Gertsch, 1 ♂ 1 ♀ ( AMNH)  ; Shasta Co., 8 km N Manzanita Lake, 19.IX.1961, W. Ivie and W.J. Gertsch, 1 ♀ ( AMNH)  ; Sierra Co., 3.2 km N Calpine, 6.IX.1959, W.J. Gertsch and V. Roth, 1 ♀ ( AMNH)  ; Sierra Co., Gold Lake , 7.VII.1952, M. Cazier, W.J. Gertsch, and R. Schrammel, 1 ♂ ( AMNH)  ; Tehama Co., Deer Creek , 19.IX.1961, W. Ivie and W.J. Gertsch, 1 ♀ ( AMNH)  ; Yuba Co., Strawberry , 12.VII.1952, M. Cazier, W.J. Gertsch, and R. Schrammel, 3 ♂ 3 ♀ ( AMNH)  ; Mountain Lassen Park , 40°N, 121°W, 28.VI.1940, W.M. Pearce, 1 ♂ ( AMNH)  ; Lassen National Park, Summit Lake , 2042 m, B. Malkin, 1.X.1944, 1 ♀ ( AMNH)  ; Sequoia National Park, Giant Forest to Marble Fork trail, 1981–2134 m, 24.VII.1907, J.C. Bradley, 1 ♂ ( AMNH)  ; Sequoia National Park , 1676 m, 5.VII.1956, V. Roth and W.J. Gertsch, 1 ♀ ( AMNH)  . Oregon: Jefferson Co., Spring Creek , 19.IV.1952, V. Roth, 1 ♂ ( AMNH)  ; Klamath Co., Lake of the Woods, 1554 m, no date data, B. Malkin, 1 ♀ ( AMNH)  ; Lake of the Woods, 1–4.VII.1934, Lawrence, 1 ♀ ( AMNH), McLoughlin Mountain base, 42°N, 122°W, 22.VI.1934, Lawrence, 1 ♂ 1 ♀ ( AMNH)  ; same data, 1 ♂ 3 ♀ ( AMNH)  ; Idle Park, North Umpqua River , 23.VIII.1959, V. Roth and W.J. Gertsch, 1 ♀ ( AMNH)  ; Crater Lake National Park, Lake Trail , 2.VIII.1951, D. Lowrie, 1 ♂ 1 ♀ ( AMNH)  ; Crater Lake National Park, 1981 m, 27.VIII.1950, B. Malkin, 1 ♀ ( AMNH)  ; 16.1 km NW Baker , 11.VII.1953, Roth and Beer, 1 ♀ ( AMNH)  ; Triangle Lake , 16.VII.1952, B. Malkin, 1 ♀ ( AMNH)  . Washington: Yakima Park , 20.VIII.1947, G. Gulion, 1 ♂ ( AMNH)  ; Seattle, 46°N, 122°W, no date and collector data, 1 ♀ (AMNH); Seattle, 46°N, 122°W, no date and collector data, 1 ♀ (AMNH).

Distribution. CANADA: British Columbia ( Bennett et al. 2014); USA: Arizona, California, Oregon, Washington (Map 1).

AMNH

American Museum of Natural History

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Agelenidae

Genus

Novalena

Loc

Novalena intermedia (Chamberlin & Gertsch 1930)

Maya-Morales, Julieta & Jiménez, María Luisa 2017
2017
Loc

Agelena intermedia

Roth 1972: 46
Lehtinen 1967: 446
Chamberlin 1942: 225
Chamberlin 1942: 226
Chamberlin 1941: 9
Loc

Novalena pima

Roth 1972: 46