Novalena creel, Maya-Morales, Julieta & Jiménez, María Luisa, 2017

Maya-Morales, Julieta & Jiménez, María Luisa, 2017, Revision of the funnel-web spider genus Novalena (Araneae: Agelenidae), Zootaxa 4262 (1), pp. 1-88 : 42-43

Novalena creel new species

Figure 20 d–h; Map 3

Type. Holotype male: MEXICO: Chihuahua: Municipality of Creel , Creel , 27°45′N, 107°38′W, 2400 m, IX.1969, V. Roth (CASENT 9048877), deposited at CAS .

Etymology. The specific name is a noun in apposition taken from the type locality.

Diagnosis. The males of N. creel differ from N. approximata and N. bosencheve in having the tegular median process with one projection ( Fig. 20 d), and from N. clara in having the RTA without a denticle on ventral projection ( Fig. 20 e). The females differ from N. approximata in having the atrium with spurs in anterior position, from N. costata and N. clara in having the atrium with straight lateral margins ( Fig. 20 f), from N. divisadero in having the primary spermathecae oblique ( Fig. 20 g), and from N. durango in having the atrium occupying less than half the plate length ( Fig. 20 f).

Description. Male (holotype): Carapace yellow, white feathery scales on cephalic region, behind eyes. Chelicerae reddish. Endites brown. Labium brown with white tip. Sternum brown with a median yellow band. Legs orange with three incomplete rings on femur, two on tibia, and one distal on metatarsus. Opisthosoma brown with yellow foliage. PLS with basal segment light brown. Total length 8.13. Carapace length 4.5, width 2.75, cephalic region width 1.52, ocular region width 0.95. Eye diameter: AME 0.13, ALE 0.21, PME 0.19, PLE 0.15. Distance between eyes: AME-AME 0.06, AME-ALE 0.1, AME-PME 0.15, ALE-PLE 0.08, ALE-ALE 0.33, PME-PME 0.15, PME-PLE 0.15. Clypeus height 0.27. Chelicerae with three retromarginal teeth; basal segment length 1.67, fang length 0.76. Labium wider than long (0.57/0.48). Endites slightly convergent (distance at their base compared at their tips 0.57/0.33). Sternum longer than wide (1.9/1.62). Opisthosoma longer than wide (4.38/2.5). ALS separated by less their basal diameter (0.24/0.36), PLS with distal segment shorter than basal segment (0.52/0.61). Leg lengths: I—femur 3.25/ patella-tibia 4/ metatarsus 3.25/ tarsus 2; II—3.5/ 4/ 3.5/ 1.88; III—3.25/ 3.88/ 3.63/ 1.88; IV—3.75/ 4.88/ 5/ 2.25. Spination: femur dorsal I—1-2 -3/ II—1-2 -3/ III—2-2 -3/ IV—1-2 -3; patella Idorsal 1-1/ prolateral 1/ retrolateral 0; II—1-1 / 1/ 0; III—1-1 / 1/ 1; IV—1-1 / 1/ 1; tibia I—dorsal 1-0-0/ ventral 2-2- 2/ prolateral 1-1-0/ retrolateral 0; II—1-1 -0/ 2-2-2/ 1-1-0/ 0; III—1-1 -0/ 2-2-2/ 1-1-0/ 1-1-0; IV—1-1 -0/ 1-1-1-2/ 1- 1-0/ 1-1-0; metatarsus I—0/ 2-2-2/ 0-1-1/ 0-0-1; II—1 -0-1/ 2-2-2/ 1-0-1/ 0-1-1; III—2-2 -2/ 2-2-2/ 0-1-1/ 0-1-1; IV—2-2 -2/ 2-2-2/ 1-1-1/ 0-0-1. Number of trichobothria on tarsus: I—6, II—7, III—5, IV—6. Number of dorsal spines on pedipalp: femur 3, tibia 4. Cymbium length 1.52, width 0.9. Embolus straight; conductor distally notched; tegular median process with rounded tip; distal projection of RTA rounded, dorsal and ventral projections of similar size ( Fig. 20 d, e).

Female (paratype) (CASENT 9048878): Carapace yellow. Chelicerae reddish. Endites orange. Labium brown with white tip. Sternum brown with a median yellow band and two pairs of lateral yellow dots. Legs orange with rings as in male, with black dots between the first two proximal rings in ventral part of femur. Opisthosoma as in male but darker. Spinnerets orange. Total length 11.25. Carapace length 5.63, width 3.63, cephalic region width 2.29, ocular region width 1.19. Eye diameter: AME 0.13, ALE 0.23, PME and PLE 0.19. Distance between eyes: AME-AME 0.19, AME-ALE 0.15, AME-PME 0.27, ALE-PLE 0.13, ALE-ALE 0.6, PME-PME 0.25, PME-PLE 0.27. Clypeus height 0.23. Chelicerae with three retromarginal teeth; basal segment length 2.27, fang length 0.95. Labium wider than long (0.82/0.7). Endites convergent (distance at their base compared at their tips 0.82/0.36). Sternum longer than wide (2.71/2.14). Opisthosoma longer than wide (6/3.88). ALS separated by less their basal diameter (0.38/0.44), PLS with distal segment slightly longer than basal segment (0.76/0.61). Leg lengths: Ifemur 4/ patella-tibia 5/ metatarsus 3.38/ tarsus 1.75; II—4 / 4.63/ 3.5/ 2.13; III—3.75/ 4.5/ 4/ 2; IV—4.75/ 5.63/ 5.63/ 2.25. Spination: femur dorsal I—1-2 -3/ II—1-3 -3/ III—1-3 -3/ IV—1-2 -3; patella I—dorsal 1-1/ prolateral 1/ retrolateral 0; II—1-1 / 1/ 0; III—1-1 / 1/ 1; IV—1-1 / 1/ 1; tibia I—dorsal 0/ ventral 2-2-2/ prolateral 1-1-0/ retrolateral 0; II—1-1 -0/ 2-2-2/ 1-1-0/ 0; III—1-1 -0/ 2-2-2/ 0-1-1/ 1-1-1; IV—1 -0-0/ 2-2-2/ 1-1-0/ 1-1-0; metatarsus I—0/ 2-2-2/ 1-1-1/ 0-0-1; II—0-0-2/ 2-2-2/ 0-1-1/ 0-0-1; III—2-1 -2/ 2-2-2/ 0-1-1/ 0-1-1; IV—2-2 -2/ 1- 1-1-2-2/ 1-0-1/ 0-0-1. Number of trichobothria on tarsus: I—7, II—7, III—7, IV—8. Dorsal spines on pedipalp femur: 3. Prolateral spines on tibia: 1-2. Epigynal plate wider than long (1.73/0.88); atrium superficial with strongly sclerotized vertical margins; anterior sclerotized spurs separated by more their basal width ( Fig. 20 f, h). Internal genitalia: copulatory ducts straight, primary spermathecae adjacent and oblique ( Fig. 20 g).

Variation. Females (n=3): body length 9.75–12.63; carapace length 5.38–5.63; patella-tibia I length 5–5.13. Spots more evident in paratype.

Additional material examined. Paratypes: MEXICO : Chihuahua: same locality from holotype, in pines, 1.XI.1973, V. Roth, 1 ♀ (CASENT 9048878) ; same data, 1 ♀ (CASENT 9048876) ; 1.VI.1991, R.E. Stecker, 1 ♀ (CASENT 9048875).

Distribution. MEXICO: Chihuahua (Map 3).

Figures

FIGURE 20. Novalena spp. genitalia: a – c, Novalena comaltepec sp. nov.; d – h, Novalena creel sp. nov. a, f, epigynum, ventral view; b, g, internal genitalia, dorsal view; c, h, epigynum, posterior view; d, male pedipalp, ventral view; e, same, retrolateral view. Abbreviations: A, atrium; C, conductor; CD, copulatory ducts; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: c, h = 0.25 mm; d = 0.5 mm.