Novalena oaxaca, Maya-Morales, Julieta & Jiménez, María Luisa, 2017

Maya-Morales, Julieta & Jiménez, María Luisa, 2017, Revision of the funnel-web spider genus Novalena (Araneae: Agelenidae), Zootaxa 4262 (1), pp. 1-88 : 60-61

Novalena oaxaca new species

Figure 32 ; Map 3

Type. Holotype male: MEXICO: Oaxaca : La Cumbre , Highway Oaxaca-Tuxtepec, 42 km N Oaxaca, 2800 m , 16.IX.2000, E. González, O. Delgado, J. Castelo, E. Lira, J. Lira, C. Durán, O. Grosselet, and M. Grosselet (CNAN-T1019), deposited at CNAN .

Etymology. The specific name is a noun in apposition taken from the state of the type locality.

Diagnosis. The males of N. oaxaca differ from N. gibarrai in having the tegular median process straight ( Fig. 32 a), and from N. perote by the absence of a ventral projection on the RTA ( Fig. 32 b). The females differ from N. attenuata in having the posterior margin not protruded ( Fig. 32 c), from N. gibarrai in having the copulatory ducts curved in lateral direction ( Fig. 32 d), and from N. perote in having the atrium without two pairs of sclerotized sharp tips ( Fig. 32 c).

Description. Male (holotype): Carapace brown, white feathery scales on cephalic region. Chelicerae dark brown, condyles brown. Endites and labium brown with white tip. Sternum dark brown with an orange central spot. Legs orange with three rings on femur. Opisthosoma dark brown with dark brown foliage and five pairs of black spots. Spinnerets yellow. Total length 6.25. Carapace length 3.13, width 2.14, cephalic region width 1.24, ocular region width 0.67. Eye diameter: AME 0.1, ALE 0.13, PME and PLE 0.15. Distance between eyes: AME-AME 0.06, AME-ALE 0.06, AME-PME 0.15, ALE-PLE 0.04, ALE-ALE 0.25, PME-PME 0.06, PME-PLE 0.08. Clypeus height 0.21. Chelicerae with three retromarginal teeth; basal segment length 1.21, fang length 0.61. Labium wider than long (0.52/0.39). Endites convergent (distance at their base compared at their tips 0.52/0.21). Sternum longer than wide (1.67/1.36). Opisthosoma longer than wide (2.88/1.88). ALS separated by less their basal diameter (0.15/0.21), PLS with distal segment shorter than basal segment (0.24/0.33). Leg lengths: I—femur 2.38/ patella-tibia 2.86/ metatarsus 1.95/ tarsus 1.57; II—2.24/ 2.62/ 1.9/ 1.33; III—2.19/ 2.43/ 1.9/ 1.29; IV—2.62/ 3.1/ 2.86/ 1.67. Spination: Femur dorsal I—1-1 -3/ II—1-3 -3/ III—1-3 -3/ IV—1-2 -3; patella I—dorsal 1-1/ prolateral 1/ retrolateral 0; II—1-1 / 1/ 0; III—1-1 / 1/ 1; IV—1-1 / 1/ 1; tibia I—dorsal 0/ ventral 2-2-2/ prolateral 1- 1-0/ retrolateral 0; II—0/ 2-2-2/ 1-1-0/ 0; III—1-1 -0/ 2-2-2/ 1-1-0/ 1-1-0; IV—1-1 -0/ 2-2-2/ 1-1-0/ 1-1-0; metatarsus I—0/ 2-2-2/ 0-1-1/ 0-0-1; II—0-0-1/ 2-2-2/ 0-1-1/ 0; III—2-1 -2/ 2-2-2/ 0-1-1/ 0-1-1; IV—2-1 -2/ 1-1-1- 2-2/ 0-1-1/ 0-0-1. Number of trichobothria on tarsus: I—6, II—6, III—6, IV—6. Number of dorsal spines on pedipalp: femur 3, tibia 3. Cymbium length 1.09, width 0.58. Embolus slightly curved; conductor distally notched; tegular median process strongly sclerotized with flat tip; ventral projection of RTA absent ( Fig. 32 a, b).

Female (paratype) (CNAN-T1021): Carapace similar to male with two longitudinal bands darker. Chelicerae brown, condyles light brown. Endites and labium brown with white tips. Sternum dark brown with diffuse orange spots. Legs light brown with three rings on femur. Opisthosoma brown with anterior spot and five pairs of posterior black spots. Spinnerets yellow. Total length 6.25. Carapace length 3, width 2.13, cephalic region width 1.21, ocular region width 0.64. Eye diameter: AME 0.1, ALE 0.13, PME 0.12, PLE 0.15. Distance between eyes: AME-AME 0.06, AME-ALE 0.04, AME-PME 0.12, ALE-PLE 0.02, ALE-ALE 0.25, PME-PME 0.08, PME-PLE 0.06. Clypeus height 0.15. Chelicerae with three retromarginal teeth; basal segment length 1.21, fang length 0.67. Labium wider than long (0.45/0.39). Endites convergent (distance at their base compared at their tips 0.45/0.21). Sternum longer than wide (1.67/1.33). Opisthosoma longer than wide (3.33/2.1). ALS separated by slightly less their basal diameter (0.19/0.21), PLS with distal segment as long as basal segment (0.29/0.29). Leg lengths: Ifemur 2.14/ patella-tibia 2.71/ metatarsus 1.76/ tarsus 1.19; II—2 / 2.24/ 1.62/ 1.05; III—1.9/ 2.24/ 1.81/ 0.9; IV— 2.52/ 3.05/ 2.71/ 1.33. Spination: femur dorsal I—1-1 -3/ II—1-3 -3/ III—1-3 -3/ IV—1-2 -3; patella I—dorsal 1-1/ prolateral 1/ retrolateral 0; II—1-1 / 1/ 0; III—1-1 / 1/ 1; IV—1-1 / 1/ 1; tibia I—dorsal 0/ ventral 2-2-2/ prolateral 1- 1-0/ retrolateral 0; II—0/ 1-2-2/ 1-1-0/ 0; III—1-1 -0/ 1-2-2/ 1-1-0/ 1-1-0; IV—1-1 -0/ 2-2-2/ 1-1-0/ 1-1-0; metatarsus I—0/ 2-2-2/ 1-0-0/ 0-0-1; II—0-0-2/ 2-2-2/ 1-0-1/ 0; III—2-1 -2/ 2-2-2/ 0-1-1/ 0-1-1; IV—3-1 -2/ 1-1-1- 2-2/ 0-1-1/ 0-1-1. Number of trichobothria on tarsus: I—6, II—6, III—6, IV—6. Dorsal spines on pedipalp femur: 3. Prolateral spines on tibia: 1-2. Epigynal plate wider than long (0.79/0.54); atrium in shallow depression; anterior hyaline spurs separated by more their basal width ( Fig. 32 c, e). Internal genitalia: copulatory ducts curved in lateral direction, primary spermathecae adjacent ( Fig. 32 d).

Variation. Males (n=3): body length 4.75–6.88; carapace length 2.63–3.75; patella-tibia I length 2.5–3.38. Females (n=2): patella-tibia I length 2.5–2.71.

Additional material examined. MEXICO: Oaxaca : same data as holotype, 2 ♂ (CNAN-T1020) ; same data, 2 ♀ (CNAN-T1021).

Distribution. MEXICO: Oaxaca (Map 3).

Figures

FIGURE 32. Novalena oaxaca sp. nov.: a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; CD, copulatory ducts; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 0.5 mm; e = 0.25 mm.