Ischnothyreus cornuatus, Edward & Harvey, 2014

Edward, Karen L. & Harvey, Mark S., 2014, Australian Goblin Spiders Of The Genus Ischnothyreus (Araneae, Oonopidae), Bulletin of the American Museum of Natural History 2014 (389), pp. 1-144 : 74-76

Ischnothyreus cornuatus , new species Figures 43–44 , map 8

TYPES: AUSTRALIA: Queensland: Male holotype and female allotype from 21 km S Atherton, 17.45 ° S, 145.4667 ° E, 1070 m (5 Nov. 1983, D. Yeates, G. Thompson), deposited in QM (³ holotype: QM S16023 , PBI _ OON 00025944 ; ♀ allotype: QM S16023 , PBI _ OON 00025757 ) .

ETYMOLOGY: The specific epithet is of the Latin cornuatus meaning ‘‘hornlike’’ ( Brown, 1956), and refers to the distal hornlike processes on the male palp.

DIAGNOSIS: The male of this species exhibits a striking modification of the embolic region of the male pedipalp, with one larger and one smaller hornlike process directed dorsally (fig. 44A, B). The female epigynal region has paired cup-shaped epigynal atriums that are large and possess thicker sclerotization at the lateral edges (figs. 43I, 44F).

MALE (PBI_OON 25944, figs. 43A–C, G– H, 44A–E). Total length 1.50. CEPHALO- THORAX: Carapace orange, broadly oval in dorsal view, pars cephalica strongly elevated in lateral view, anteriorly narrowed to 0.49 times its maximum width or less, anterolateral corners with slightly sclerotized triangular projections, surface of elevated portion of pars cephalica finely reticulate, sides strongly reticulate. Clypeus margin unmodified, straight in front view, low, ALE separated from edge of carapace by less than their radius; setae dark. Eyes: ALE largest, ALE circular, PME oval, PLE oval; posterior eye row straight from above; ALE touching, ALE-PLE touching. Sternum longer than wide, pale orange, uniform; setae dark, evenly scattered. Chelicerae, endites, and labium pale orange. Chelicerae slightly divergent, anterior face unmodified; promargin with one larger denticle; fang normal, without prominent basal process. Labium elongated hexagon, not fused to sternum, anterior margin indented at middle, slightly stronger sclerotization at base; with six or more setae on anterior margin. Endites anteromedian tip with one strong, toothlike projection, slightly stronger sclerotization than sternum. ABDO- MEN: Ovoid; dorsum soft portions white. Book lung covers elliptical. Dorsal scutum pale orange, covering more than 3/4 of abdomen, more than 1/2 to most of abdomen width, middle surface smooth, sides smooth. Epigastric scutum small lateral sclerites absent. Postepigastric scutum pale orange, covering about 1/2 of abdominal length. Dorsum, epigastric area, and postepigastric area setae dark. LEGS: Yellow, without color pattern; patella plus tibia I shorter than carapace. Leg spination: femora: I p0-2-0; II p0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Palp proximal segments dark red-brown; embolus dark; femur shorter than trochanter; patella shorter than femur; cymbium dark red-brown; bulb dark red-brown, more than 2 times as long as cymbium, elongate; embolic region with two hornlike processes, one large and one small, both directed dorsally (fig. 44A, B).

FEMALE (PBI_OON 25757, figs. 43D–F, I, 44F). Total length 1.73. ABDOMEN: Dorsal scutum covering 1/2 to 3/4 of abdomen, between 1/4 and 1/2 abdomen width. Epigastric scutum small lateral sclerites present. Postepigastric scutum widely hexagonal, covering about 1/3 of the abdominal length. LEGS: Pale orange. Leg spination: femora: I p0-2-0; II p0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Ventral view: epigynal atrium paired, cup shaped, sclerotization heavier at lateral edges; convoluted duct very long, thicker than apodemes, ending posteriorly at small cresent-shaped indent (figs. 43I, 44F).

OTHER MATERIAL EXAMINED: AUS- TRALIA: Queensland: 21 km S Atherton , rainforest, leaf litter, 1070 m, 17.45000 ° S, 145.46670 ° E, Nov. 5, 1983 (D. Yeates, G. Thompson, QM S16022 , PBI _OON 25797), 1 ³ ; Millaa Millaa Lookout , rainforest, 1000 m, 17.51667 ° S, 145.56670 ° E, Dec. 1, 1993, to Feb. 25, 1994 (J. Hasenpusch, QM S49694 , PBI _OON 22107), 1 ³ ; Mount Fisher , 1150 m, 17.33000 ° S, 145.32000 ° E, May 3–June 2, 1995 (P. Zborowski, ANIC, PBI _ OON 5847 ), 1 ♀ ; Mount Fisher ( Kjelberg ), rainforest, leaf litter, 1100 m, 17.53333 ° S, 145.55000 ° E, May 18, 1995 (G. Monteith, QM S41902 , PBI _OON 21955), 1 ♀ ; Mount Fisher, 7 km SW. of Millaa Millaa , rainforest, leaf litter, 1100 m, 17.55000 ° S, 145.55000 ° E, Apr. 27, 1982 (G. Monteith, D. Yeates, D. Cook, QM S12995 , PBI _OON 25799), 1 ³ ; Tower nr the Crater , rainforest, leaf litter, 1230 m, 17.45000 ° S, 145.48330 ° E, Nov. 23, 1994 (G. Monteith, QM S31611 , PBI _OON 22177), 1 ♀ ; Tower nr the Crater National Park , rainforest, 1230 m, 17.45000 ° S, 145.48330 ° E, Nov. 25, 1994, to Jan. 10, 1995 (G. Monteith, J. Hasenpusch, QM S46987 , PBI _OON 21982), 3 ³, 2 ♀ .

DISTRIBUTION: This species is known only from the Atherton Tableland (AU subregion), of the Wet Tropics Bioregion, in northeastern Queensland (map 8).

Figures

Fig. 43. Ischnothyreus cornuatus, sp. nov. Holotype male (PBI_OON 00025944): A. Habitus, dorsal view; B. habitus, ventral view; C. habitus, lateral view; G. carapace, anterior view; H. sternum, ventral view. Allotype female (PBI_OON 00025757): D. habitus, dorsal view; E. habitus, ventral view; F. habitus, lateral view; I. epigynum, ventral view.

Fig. 44. Ischnothyreus cornuatus, sp. nov. Holotype male (PBI_OON 00025944): A. left palp, prolateral view; B. left palp, retrolateral view; C. left chelicerae, anterior view; D. left chelicerae, posterior view; E. left palp, dorsal view. Allotype female (PBI_OON 00025757): F. epigynum, ventral view. Scale lines 5 0.1 mm.