Ischnothyreus corniculatum, Edward & Harvey, 2014

Edward, Karen L. & Harvey, Mark S., 2014, Australian Goblin Spiders Of The Genus Ischnothyreus (Araneae, Oonopidae), Bulletin of the American Museum of Natural History 2014 (389), pp. 1-144 : 77-81

Ischnothyreus corniculatum , new species Figures 45–46 , map 6

TYPES: AUSTRALIA: Queensland: Male holotype, female allotype and three female paratypes from Herberton Range State Forest, Forestry track, 17.27111 ° S, 145.42277 ° E (23 Apr. 2009, K. Edward and P. Cullen), deposited in QM (³ holotype: QM S95945 , PBI _ OON 00005611 ; ♀ allotype and paratypes: QM S95946 , PBI _ OON 00026309 ) .

ETYMOLOGY: The specific epithet is the Latin corniculatum meaning ‘‘little horn’’ ( Brown, 1956), and refers to the small hornshaped process on the distal tip of the male palp.

DIAGNOSIS: Like I. cornuatus , sp. nov., the male of this species possesses a hornlike modification on the embolic region of the male pedipalp. However, I. corniculatum , sp. nov., has two small lateral processes and only one dorsal hornlike process, instead of two (fig. 46 A, B). The epigynal atrium of the female epigynal region is not paired, and exists as a wide, but short depression that is heavily sclerotized around the posterior edge (figs. 45J, 46F).

MALE (PBI_OON 5611, figs. 45A–C, G–I, 46A–E). Total length 1.53. CEPHALOTHO- RAX: Carapace yellow-brown, broadly oval in dorsal view, pars cephalica strongly elevated in lateral view, anteriorly narrowed to between 0.5 and 0.75 times its maximum width, anterolateral corners without extension or projections, surface of elevated portion of pars cephalica strongly reticulate, sides strongly reticulate. Clypeus margin unmodified, curved downward in front view, high, ALE separated from edge of carapace by their radius or more; setae dark. Eyes: ALE largest, ALE oval, PME circular, PLE oval; posterior eye row straight from above; ALE touching, ALE-PLE touching. Sternum as long as wide, pale orange, darker anteriorly near labium forming unique pattern (fig. 45H, I); setae dark, evenly scattered. Chelicerae, endites, and labium dark redbrown. Chelicerae straight, anterior face unmodified; promargin with one larger denticle; fang shape normal, without prominent basal process; setae dark. Labium elongated hexagon, not fused to sternum, anterior margin indented at middle, much more heavily sclerotized than sternum, with six or more setae on anterior margin. Endites anteromedian tip with one strong, toothlike projection, much more heavily sclerotized than sternum. ABDOMEN: Ovoid; dorsum soft portions white. Book lung covers elliptical. Dorsal scutum yellow-brown, covering 1/2 to 3/4 of abdomen, more than 1/2 to most of abdomen width, middle surface finely reticulate, sides finely reticulate. Epigastric scutum small lateral sclerites present. Postepigastric scutum pale orange, covering about 1/2 of abdominal length. Dorsum, epigastric area, and postepigastric area setae dark. LEGS: Pale orange, without color pattern; patella plus tibia I shorter than carapace. Leg spination: femora: I p0-2-0; II p0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Palp proximal segments dark red-brown; embolus dark; femur shorter than trochanter; patella about as long as femur; cymbium dark redbrown; bulb dark red-brown, more than two times as long as cymbium, stout; embolic region with one posteriorly directed hornlike process and two small lateral processes (fig. 46A, B).

FEMALE (PBI_OON 26309, figs. 45D–F, J, 46F). Total length 1.50. CEPHALOTHO- RAX: Carapace pars cephalica slightly elevated in lateral view, anteriorly narrowed to 0.49 times its maximum width or less. Chelicerae, endites, and labium pale orange. ABDOMEN: Dorsal scutum between 1/4 and 1/2 abdomen width. Postepigastric scutum widely hexagonal, covering about 1/3 of the abdominal length. LEGS: Leg spination: femora: I p0-2-0; r0-1-1; II p0-1-0; r0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1- 1-0; v1-1-0; spines on retrolateral side much smaller than prolateral side. GENITALIA: Ventral view: epigynal atrium wider than high, sclerotization heavier around curved posterior edge; convoluted duct thicker than apodemes (figs. 45J, 46F).

OTHER MATERIAL EXAMINED: AUS- TRALIA: Queensland: 2 km E of South Johnstone Forestry Camp , 17.60000 ° S, 146.00000 ° E, Dec. 1, 1993, to Feb. 25, 1994 (J. Hasenpusch, QM S63216 , PBI _OON 22037), 1 ♀ ; Boulder Creek, via Tully , rainforest, leaf litter, 17.83333 ° S, 145.90000 ° E, Oct. 27, 1983 (G. Monteith, D. Yeates, G. Thompson, QM S16024 , PBI _OON 25798), 1 ³ ; Downey Creek , 25 km SE Millaa Millaa, 400 m, 17.65000 ° S, 145.78330 ° E, Dec. 7, 1988 (G. Monteith, G. Thompson, QM S31962 , PBI _OON 21746), 1 ♀ ; Herberton Range State Forest, Forestry track, rainforest, leaf litter, 17.27111 ° S, 145.42277 ° E, Apr. 23, 2009 (K. Edward and P. Cullen, QM S95947 , PBI _ OON 5535 ), 1 ³ ; Kenny Road , 850 m, 17.46667 ° S, 145.53330 ° E, Nov. 25, 1994, to Jan. 10, 1995 (G. Monteith, J. Hasenpusch, QM S27980 , PBI _OON 21789), 1 ³ ; Longlands Gap SF, forest, 17.45500 ° S, 145.47420 ° E, July 23–Nov. 26, 1992 (R. Raven, P. Lawless, E. Lawless, M. Shaw, QM S19940 , PBI _OON 25927), 1 ³, 1 ♀ ; Malaan SF, rainforest, 17.58834 ° S, 145.60750 ° E, Nov. 26, 1992, to Apr. 15, 1993 (R. Raven, P. Lawless, QM S24383 , PBI _OON 21735), 1 ³ ; Malaan SF, rainforest, 1000 m, 17.58333 ° S, 145.58330 ° E, Apr. 20–24, 1978 (R. Raven, V. Davies, QM S16126 , PBI _OON 25796), 4 ³ ; Malaan SF, rainforest, 1000 m, 17.58333 ° S, 145.58330 ° E, Apr. 20–24, 1978 (R. Raven, V. Davies, QM S16126 , PBI _OON 25915), 3 ³ ; McNamee Creek , rainforest, leaf litter, 400 m, 17.63333 ° S, 145.8500 ° E, July 8, 1971 (Taylor and Feehan, ANIC, PBI _OON 25848), 2 ³ ; Mount Fisher, 7 km SW Millaa Millaa, Kjellberg Road , 1000 m, 17.55000 ° S, 145.56670 ° E, May 3, 1983 (G. Monteith, D. Yeates, QM S16097 , PBI _OON 25800), 1 ³ ; Mount Fisher, 7 km SW Millaa Millaa, Kjellberg Road , 1000 m, 17.55000 ° S, 145.56670 ° E, May 3, 1983 (G. Monteith, D. Yeates, QM S16091 , PBI _OON 26241), 2 ♀ ; Mount Fisher, Kjellberg Road , 1000 m, 17.53333 ° S, 145.55000 ° E, Dec. 1, 1993, to Feb. 25, 1994 (J. Hasenpusch, QM S34940 , PBI _OON 21947), 1 ³ ; Palmerston National Park , rainforest, 670 m, 17.58834 ° S, 145.70000 ° E, Nov. 30, 1992, to Apr. 15, 1993 (R.J. and S. Raven, P. and E. Lawless, QM S22948 , PBI _OON 25919), 1 ³ ; Palmerston National Park , rainforest, 670 m, 17.58834 ° S, 145.70000 ° E, July 25–Nov. 30, 1992 (R. Raven, P. Lawless, E. Lawless, M. Shaw, QM S21922 , PBI _OON 25931), 1 ³ ; Palmerston National Park, Downey Creek Road, rainforest, 17.60500 ° S, 145.76670 ° E, July 25–Nov. 30, 1992 (R. Raven, P. Lawless, M. Shaw, QM S24238 , PBI _OON 21705), 2 ³ ; Palmerston National Park, Downey Creek Road, rainforest, 17.60500 ° S, 145.76670 ° E, Oct. 30, 1991, to July 24, 1992 (R. Raven, P. Lawless, M. Shaw, QM S24577 , PBI _OON 21708), 2 ♀ ; the Millstream , 10 km NNE Ravenshoe, 1040 m, 17.53333 ° S, 145.51670 ° E, Dec. 6, 1998, to Feb. 5, 1999 (G. Monteith, D. Cook, QM S49225 , PBI _OON 22202), 1 ³, 1 ♀ ; Tully Gorge National Park , rainforest, leaf litter, 750 m, 17.7000 ° S, 145.5500 ° E, July 2, 1971 (Taylor and Feehan, ANIC, PBI _ OON 5848 ), 1 ♀ ; Upper Boulder Creek , 8 km N of Tully, 250 m, Dec. 4–7, 1989 (G. Monteith, G. Thompson, H. Janetzki, QM S46824, PBI _OON 22027), 1 ♀ ; Upper Boulder Creek, Walter Hill Range , 950 m, 17.83333 ° S, 145.90000 ° E, Nov. 17–18, 1984 (V. Davies, G. Monteith, J. Gallon, D. Cook, G. Thompson, QM S12923 , PBI _ OON 6434 ), 1 ♀ ; Wooroonooran National Park, Tchupala Falls track, rainforest, leaf litter, 17.61000 ° S, 145.77722 ° E, Apr. 27, 2009 (K. Edward and P. Cullen, QM S95948 , PBI _ OON 5566 ), 1 ♀ ; Wooroonooran National Park, Tchupala Falls track, rainforest, leaf litter, 17.61000 ° S, 145.77722 ° E, Apr. 27, 2009 (K. Edward and P. Cullen, QM S95949 , PBI _OON 26306), 1 ♀ ; Wooroonooran National Park, Tchupala Falls track, rainforest, leaf litter, 17.61000 ° S, 145.77722 ° E, May 17, 2007 (K. Edward and K. Pitz, QM S95950 , PBI _OON 26312), 1 ♀ .

DISTRIBUTION: This species is only known from the Atherton Tablelands (AU) and the Cairns-Cardwell Lowlands (CC), of the Wet Tropics Bioregion, in northeastern Queensland (map 6).

Figures

Fig. 45. Ischnothyreus corniculatum, sp. nov. Holotype male (PBI_OON 00005611): A. habitus, dorsal view; B. habitus, ventral view; C. habitus, lateral view; G. carapace, anterior view; H. sternum, ventral view; I. sternum, oblique view. Allotype female (PBI_OON 00026309): D. habitus, dorsal view; E. habitus, ventral view; F. habitus, lateral view; J. epigynum, ventral view.

Fig. 46. Ischnothyreus corniculatum, sp. nov. Holotype male (PBI_OON 00005611): A. left palp, prolateral view; B. left palp, retrolateral view; C. left chelicerae, anterior view; D. left chelicerae, posterior view; E. left palp, dorsal view. Allotype female (PBI_OON 00026309): F. epigynum, ventral view. Scale lines 5 0.1 mm.