Ischnothyreus eacham, Edward & Harvey, 2014

Edward, Karen L. & Harvey, Mark S., 2014, Australian Goblin Spiders Of The Genus Ischnothyreus (Araneae, Oonopidae), Bulletin of the American Museum of Natural History 2014 (389), pp. 1-144 : 84-87

Ischnothyreus eacham , new species Figures 49–50 , map 8

TYPES: AUSTRALIA: Queensland: Male holotype, female allotype, and two female paratypes from Eacham National Park , 17.2833 ° S, 145.6167 ° E, 760 m (20 Feb 1973, R.W. Taylor), deposited in ANIC (³ holotype: PBI _ OON 00005883 ; ♀ allotype and paratypes: PBI _ OON 00005854 ) .

ETYMOLOGY: The specific epithet is a noun in apposition, taken from the type locality.

DIAGNOSIS: The male of this species exhibits a simplified male palp, with a broad, obtusely bent distal tip. A small lateral process bends backward on the retrolateral side of the embolic region (fig. 50A, B, E). The female epigynal region is equally simple, and consists of a small triangular extention pointing anteriorly and positioned at the base of a long convoluted duct (figs. 49I, 50F).

MALE (PBI_OON 5883, figs. 49A–C, G– H, 50A–E). Total length 1.63. CEPHALO- THORAX: Carapace orange-brown, broadly oval in dorsal view, pars cephalica strongly elevated in lateral view, anteriorly narrowed to 0.49 times its maximum width or less, anterolateral corners without extension or projections, surface of elevated portion of pars cephalica smooth, sides finely reticulate. Clypeus margin unmodified, curved downward in front view, low, ALE separated from edge of carapace by less than their radius; setae dark. Eyes: ALE largest, ALE circular, PME oval, PLE oval; posterior eye row straight from above; ALE touching, ALE- PLE touching. Sternum longer than wide, pale orange, uniform; setae dark, evenly scattered. Labium and endites dark red brown, chelicerae pale. Chelicerae slightly divergent, anterior face unmodified; promargin with one or two slightly larger denticles; fang shape normal, without prominent basal process; setae dark. Labium elongated hexagon, not fused to sternum, anterior margin indented at middle, same as sternum in sclerotization; with six or more setae on anterior margin. Endites anteromedian tip with one strong, toothlike projection, much more heavily sclerotized than sternum. AB- DOMEN: Ovoid; dorsum soft portions white. Book lung covers elliptical. Dorsal scutum pale orange, covering 1/2 to 3/4 of abdomen, more than 1/2 to most of abdomen width, middle surface smooth, sides smooth. Epigastric scutum small lateral sclerites present. Postepigastric scutum pale orange, covering about 2/3 of abdominal length. Dorsum, epigastric area, and postepigastric area setae dark. LEGS: Pale orange, without color pattern; patella plus tibia I shorter than carapace. Leg spination: femora: I p0-1-1; II p0-2-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Palp proximal segments dark red-brown; embolus dark; femur one to two times as long as trochanter; patella about as long as femur; cymbium dark red-brown; bulb dark red-brown, 1 to 1.5 times as long as cymbium, stout; embolic region stout, broad, obtusely bent, small lateral process bent backward on retrolateral side (fig. 50A, B, E).

FEMALE (PBI_OON 00005854, figs. 49D– F, I, 50F). Total length 1.90. CEPHALO- THORAX: Carapace pale orange, pars cephalica slightly elevated in lateral view, surface of elevated portion of pars cephalica finely reticulate. Clypeus straight in front view. Chelicerae, endites, and labium pale orange. ABDOMEN: Dorsal scutum between 1/4 and 1/2 abdomen width. Postepigastric scutum widely hexagonal, only around epigastric furrow. LEGS: Leg spination: femora: I p0-1-1; II p0-2-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Ventral view: epigynal region simple, epigynal atrium not clearly visible but with slight depression, posterior portion of depression with small triangular extention pointing anteriorly; convoluted duct thicker than apodemes, long (figs. 49I, 50F).

OTHER MATERIAL EXAMINED: AUS- TRALIA: Queensland: Curtain Fig Tree , 1 km SW of Yungaburra, rainforest, leaf litter, 17.27472 ° S, 145.57333 ° E, Dec. 3, 1969 (J.G. Brooks, ANIC, PBI _ OON 5839 ), 1 ♀ ; 1

♀ ( ANIC PBI _ OON 5839 ) ; Danbulla National Park , Mobo Creek crater circuit walk, rainforest, leaf litter, 17.16944 ° S, 145.63888 ° E, Apr. 22, 2009 (K. Edward and P. Cullen, QM S95953 , PBI _OON 26316), 1 ³ ; Dinden National Park , start of Kahlpahlim rock trail, rainforest, leaf litter, 17.03388 ° S, 145.61305 ° E, Apr. 11, 2009 (K. Edward and J. Waldock, QM S95954 ), 3 ♀ ; Dinden National Park , start of Kahlpahlim rock trail, rainforest, leaf litter, 17.03388 ° S, 145.61305 ° E, Apr. 11, 2009 (K. Edward and J. Waldock, QM S95955 , PBI _OON 26318), 3 ♀ ; Boar Pocket Road , 5 miles N. of Gillies highway, rainforest, leaf litter, 17.20000 ° S, 145.66666 ° E, Dec. 13, 1969 (J.G. Brooks, ANIC, PBI _OON 25850), 1 ³, 2 ♀ ; Boar Pocket Road , 5 miles N. of Gillies highway, rainforest, leaf mould, 17.20000 ° S, 145.66666 ° E, Nov. 15, 1969 (J.G. Brooks, ANIC, PBI _OON 25872), 1 ♀ ; Cathedral Tree , 720 m, 17.18333 ° S, 145.65000 ° E, Feb. 7, 1996 (G. Monteith, QM S43828 , PBI _OON 22053), 1 ♀ ; Crater Lakes National Park , Lake Barine walk trail, rainforest, leaf litter, 17.23722 ° S, 145.64083 ° E, Apr. 23, 2009 (K. Edward and P. Cullen, QM S95956 , PBI _ OON 5562 ), 1 ♀ ; Crater Lakes National Park , Lake Eacham walk, rainforest, leaf litter, 17.28805 ° S, 145.62166 ° E, May 18, 2007 (K. Edward and K. Pitz, QM S95957 ), 1 ♀ ; Crater Lakes National Park , Lake Eacham walk, rainforest, leaf litter, 17.28805 ° S, 145.62166 ° E, May 18, 2007 (K. Edward and K. Pitz, QM S95958 , PBI _OON 25719), 2 ♀ ; Crater Lakes National Park , Lake Eacham walk, rainforest, leaf litter, 17.28805 ° S, 145.62166 ° E, May 18, 2007 (K. Edward and K. Pitz, QM S95959 , PBI _OON 25720), 1 ³ ; Davies Creek Road, 20 km ESE Mareeba, 750 m, Dec. 4–13, 1988 (G. Monteith, G. Thompson, QM S49364, PBI _OON 22080), 1 ♀ ; Eacham National Park , rainforest, leaf litter, 760 m, 17.2833 ° S, 145.6167 ° E, Oct. 1–7, 1972 (R.W. Taylor, ANIC, PBI _ OON 5842 , 5854 ), 4 ♀ ; Eacham National Park , rainforest, 760 m, 17.2833 ° S, 145.6167 ° E, Feb. 16, 1973 (R.W. Taylor, ANIC, PBI _ OON 5859 ), 1 ♀ ; Eacham National Park , rainforest, 760 m, 17.2833 ° S, 145.6167 ° E, Mar. 23, 1975 (R.W. Taylor, ANIC, PBI _ OON 5862 ), 1 ³, 2 ♀ ; Eacham National Park , rainforest, leaf litter, 760 m, 17.2833 ° S, 145.6167 ° E, Oct. 1–7, 1972 (R.W. Taylor, ANIC, PBI _OON 25851), 1 ♀ ; Eacham National Park , rainforest, leaf litter, 760 m, 17.2833 ° S, 145.6167 ° E, Mar. 21, 1975 (R.W. Taylor, ANIC, PBI _OON 25854), 1 ♀ ; Gadgarra Road , 5 km E Lake Eacham, 700 m, 17.26667 ° S, 145.66670 ° E, Dec. 9, 1989 (G. Monteith, G. Thompson, H. Janetzki, QM S31926 , PBI _OON 22232), 1 ³, 2 ♀ ; Mount Edith Road , rainforest, leaf litter, 1000 m, 17.06777 ° S, 145.57305 ° E, Apr. 9, 2009 (K. Edward and J. Waldock, QM S95960 , PBI _ OON 5533 ), 1 ³, 1 ♀ ; Mount Edith Road , rainforest, leaf litter, 1000 m, 17.06777 ° S, 145.57305 ° E, Apr. 9, 2009 (K. Edward and J. Waldock, QM S95961 , PBI _OON 26317), 2 ³, 3 ♀ ; same data ( WAM T130762 ), 2 ³, 2 ♀ ; North Bell Peak , Malbon Thompson Range, 17.11667 ° S, 145.88330 ° E, Nov. 20–22, 1990 (G. Monteith, G. Thompson, QM S47016 , PBI _OON 22066), 1 ♀ ; Tolga , near Atherton, rainforest, leaf litter, 17.17805 ° S, 145.45916 ° E, June 14, 1978 (A. and M. Walford-Huggins, ANIC, PBI _OON 25806), 1 ³ ; Wongabel SF, rainforest, 17.32167 ° S, 145.49080 ° E, Nov. 05, 1991, to July 23, 1992 (R. Raven, P. Lawless, E. Lawless, M. Shaw, QM, S24209 , PBI _OON 21747), 2 ♀ ; Wongabel SF, 17.32167 ° S, 145.49080 ° E, July 23–Nov. 26, 1992 (R. Raven, P. Lawless, E. Lawless, M. Shaw, QM S22607 , PBI _OON 22098), 1 ³ ; Wongabel SF, forest walk, rainforest, under bark, 17.32500 ° S, 145.49583 ° E, Apr. 22, 2009 (K. Edward and P. Cullen, QM S95962 , PBI _ OON 5561 ), 1 ♀ ; Wooroonooran National Park , Tchupala Falls track, rainforest, leaf litter, 17.61000 ° S, 145.77722 ° E, Apr. 27, 2009 (K. Edward and P. Cullen, QM S95963 , PBI _ OON 5557 ), 1 ³ ; Wooroonooran National Park , Tchupala Falls track, rainforest, leaf litter, 17.61000 ° S, 145.77722 ° E, Apr. 27, 2009 (K. Edward and P. Cullen, QM S95964 , PBI _OON 26307), 2 ³ .

DISTRIBUTION: This species is known only from three central subregions of the Wet Tropics Bioregion (AU, LU, and MT), in northeastern Queensland (map 8).

Figures

Fig. 49. Ischnothyreus eacham, sp. nov. Holotype male (PBI_OON 00005883): A. habitus, dorsal view; B. habitus, ventral view; C. habitus, lateral view; G. carapace, anterior view; H. sternum, ventral view. Allotype female (PBI_OON 00005854): D. habitus, dorsal view; E. habitus, ventral view; F. habitus, lateral view; I. epigynum, ventral view.

Fig. 50. Ischnothyreus eacham, sp. nov. Holotype male (PBI_OON 00005883): A. left palp, prolateral view; B. left palp, retrolateral view; C. left chelicerae, anterior view; D. left chelicerae, posterior view; E. left palp, dorsal view. Allotype female (PBI_OON 00005854): F. epigynum, ventral view. Scale lines 5 0.1 mm.