Archaboilus polyneurus sp. nov. Gu, Yue & Ren

Gu, Jun-Jie, Yang, Xin, Huang, Rong, Yang, Guijun, Yue, Yanli & Ren, Dong, 2021, New species and material of Hagloidea (Insecta, Ensifera) from the Yanliao biota of China, ZooKeys 1033, pp. 183-190 : 183

Archaboilus polyneurus sp. nov. Gu, Yue & Ren Fig. 1

Diagnosis.

ScA reaches anterior wing margin at level of divergence of M+ CuA, RP branched distally, lengths of free CuA and free M equal, CuA + CuPaα with numerous branches.

Material examined.

Holotype, CNU-ORT-NN2009018PC. Paratype, CNU-ORT-NN2009011.

Locality and age.

Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China; Jiulongshan Formation, Bathonian-Callovian boundary interval ( Ren et al. 2019), Middle Jurassic.

Description.

Forewing oval, estimated length ca 33 mm. ScA crossing area between ScP and anterior wing margin, reaching margin at level of divergence of M+ CuA; basal part of ScP slightly anteriorly curved, ScP reaching anterior margin at 3/4 to wing base with numerous oblique branches uniformly distributed; branches of ScP with secondary vein between them, formed by two rows of cells; most cross-veins between ScP and R straight; stem R slightly undulate; RA basally branched, pectinate with 4-7 terminal branches; base of RP curved towards to posterior margin, RP very distally branched with less branches than RA; area between RA and RP with series of regular arranged cross-veins; area between R and M expanding when R dichotomous, with series of long cross-veins, cross-veins of expanded area curved; presence of a transverse veinlet connecting MA and base of RP (asterisk on Fig. 1B, D, F ); M separated from M + CuA distant to origin of RP; MA probably undulate; MP strongly curved basally (not preserved in holotype); lengths of free CuA and free M equal; CuA + CuPaα with numerous branches; CuPaβ oblique; “handle” straight; CuPb strongly oblique, basal part and middle part (where bearing teeth) forms obtuse angle.

Etymology.

From the Latin " polyneurus ", referring to its numerous branches of CuA + CuPaα.

Discussion.

Although the preservation and deformation of the specimens makes it difficult to identify the complete structure of ScA, this new species can be assigned to Archaboilus Martynov, 1937 by a combination of its ScA crossing the area between ScP and the anterior wing margin, the base of MP strongly curved, and the presence of a transverse veinlet connecting MA and the base of RP. Besides these diagnostic characters of the genus, A. polyneurus sp. nov. shares with A. musicus from the same locality a slightly sigmoidal ScP, but it differs from it by its much more distally branched RP and distinctly smaller forewing. Although the holotype and single known specimen of A. kisylkiensis Martynov, 1937 is only the basal half of a forewing, its free CuA is much longer than its free M, not as in the new species. A. polyneurus sp. nov. differs from all other Archaboilus species by its shorter ScA, very distally branches of RP, and numerous branches of CuA + CuPaα. Although the terminals numbers of RA and CuA + CuPaα are different between the holotype and paratype, this kind of difference has been shown to be intra-specific variation in orthopterans and their relatives ( Béthoux 2008; Gu et al. 2010, 2011).

Figures

Figure 1. Photos and drawings of Archaboilus polyneurus sp. nov. Gu, Yue & Ren, asterisk indicates the transverse veinlet connecting MA and base of RP. A-D right and left forewing of the holotype, CNU-ORT-NN 2009018 C E, F CNU-ORT-NN 2009011. Scale bar: 5 mm.