Coniceromyia circulata, Ament & Kung & Brown, 2020

Ament, Danilo César, Kung, Giar-Ann & Brown, Brian V., 2020, Forty-one new species of Coniceromyia Borgmeier (Diptera: Phoridae), an identification key, and new distributional records for the species of the genus, Zootaxa 4830 (1), pp. 1-61 : 8-9

Coniceromyia circulata sp. nov.

( Figs. 2C , 8I , 13I , 18E , 24I , 26I )

Diagnosis (male). Wing with round apical macula between M 1 and M 2, foretrochanter with ventral process bearing apical group of curved setae.

Material examined. Holotype ♂, BRAZIL: Amazonas : Tapauá : Rio Ipixuna: Porto Cotinha, 8–13.x.2013, Malaise trap, D.M.M. Mendes col. ( MZUSP) [MZ053401].

Description. Male. Body length 1.6 mm. Head. Frons brown, sparsely setulose, with well-defined median furrow. Flagellomere 1 light brown, conical-elongate. Arista apical, slightly longer than flagellomere 1. Palpus light brown. Gena with two setae. Thorax. Scutum, scutellum, and pleural regions brown; anepisternum bare. Legs yellowish-brown, except femora brown. Foretrochanter with ventral process bearing apical group of curved setae. Foretibia with two dorsal setae and anterodorsal row of strong setulae. Foremetatarsus with anterior excavation orange, anteroapical setulose process with strong long seta at its base, base of excavation with group of setulae ( Fig. 8I ). Foremetatarsus ratio 0.44. Posteroventral face of hind femur with narrow band of tiny blunt setulae ( Fig. 18E ). Hind femur height/length ratio 0.36. Hind tibia with one dorsal seta in basal half. Wing ( Fig. 13I ). Round apical macula between M 1 and M 2; Costa swollen, 0.4 of the wing length; R 2 + 3 absent; M 1 concave, curved posteriorly at apex; M 2 approximately concave, CuA 1 -M 2 /M 2 -M 1 ratio 1.03; CuA 1 sinuous. Halter yellowish-brown. Abdomen. Tergites brown. Hypopygium light brown. Left epandrial process dorsally setose, with ventral lobe, dorsal medially directed projection near hypoproct, subepandrial rounded elevation near apex and subepandrial digitiform setulose process. Margin of right lobe of hypandrium without lateral projection. Two setae on hypoproct ( Figs. 24I ).

Distribution. Known from a single site in Brazilian Amazonia.

Etymology. Name derived from the Latin word circulus, referring to the round apical macula between M 1 and M 2.

Figures

FIGURE 2. Habitus of the holotypes of new species of Coniceromyia, continuation (scale bars=1mm). A. C. camptoneura, B. C. chrysopa, C. C. circulata, D. C. cladopyga, E. C. crassivena, F. C. dasypoda.

FIGURE 8. Male foremetatarsus of new species of Coniceromyia (scale bars=0.1mm). A. C. amazonensis, B. C. anisocrossa, C. C. balbiae, D. C. bellatula, E. C. bisetosa, F. C. brachypoda, G. C. camptoneura, H. C. chrysopa, I. C. circulata, J. C. cladopyga, K. C. crassivena, L. C. dasypoda, M. C. deltopoda, N. C. diadela, O. C. diffusa, P. C. dolichopoda, Q. C. hadrochaeta, R. C. hirtipenna, S. C. inflata, T. C. leia, U. C. litopoda, V. C. luna, W. C. maculata, X. C. megalosoma, Y. C. memorialis, Z. C. niemeyeri, A’. C. numerosa, B’. C. ochyrosticha, C’. C. parallela, D’. C. pectinella.

FIGURE 13. Wings of the males of Coniceromyia new species. A. C. amazonensis, B. C. anisocrossa, C. C. balbiae, D. C. bellatula, E. C. bisetosa, F. C. brachypoda, G. C. camptoneura, H. C. chrysopa, I. C. circulata, J. C. cladopyga, K. C. crassivena, L. C. dasypoda.

FIGURE 18. Hind femora of the males of Coniceromyia new species, continuation (scale bars on figures of the entire femur=0.3 mm, on figures of ventrobasal part of the femur=0.1 mm). A–B. C. camptoneura, C–D. C. chrysopa, E. C. circulata, F–G. C. cladopyga, H–I. C. crassivena, J–K. C. dasypoda.

FIGURE 24. Hypopygia of Coniceromyia new species in right lateral view (except figure J which is left lateral view). A. C. amazonensis, B. C. anisocrossa, C. C. balbiae, D. C. bellatula, E. C. bisetosa, F. C. brachypoda, G. C. camptoneura, H. C. chrysopa, I. C. circulata, J. C. cladopyga in left lateral view, K. C. cladopyga in right lateral view, L. C. crassivena, M. C. dasypoda, N. C. deltopoda, O. C. diadela, P. C. diffusa, Q. C. dolichopoda, R. C. hadrochaeta, S. C. hirtipenna, T. C. inflata, U. C. leia, V. C. litopoda, W. C. luna, X. C. maculata.

FIGURE 26. Unique or uncommon features of males of Coniceromyia species. A. Curved forecoxa of C. broadheadi, B–C, Sculpturing in dorsum of forefemur of C. broadheadi, D. Anterior view of foretibia of C. leia, showing the ventral face mostly bare and shiny but with differentiated thin setulae, E. Hind tibia of C. piricornis with dorsal patch of differentiated setulae, F. Hind tibia of C. diadella with dorsal carina with differentiated setulae, G. Posteroventral face of forefemur of C. systenopoda with a triangular ventral projection, H. Forefemur of C. pectinella with dorsal excavation bearing a comb of apically-directed curved setae, I. Foretrochanter of C. circulata with ventral process bearing apical group of curved setae, J. Foretrochanter of C. bellatula with ventral projection bearing apical group of curved setulae, K. Forefemur of C. chrysopa with bare oval ring surrounding an oval patch with small setulae, L. Forefemur of C. solaris with bare oval ring surrounding an oval patch with small setulae, M. Anterior face of midfemur of C. piricornis with ventral patch of dense setulae, N. Anterior face of midfemur of C. setosa with band of tiny setulae along basal two-thirds of ventral margin.