Coniceromyia diadella, Ament & Kung & Brown, 2020

Ament, Danilo César, Kung, Giar-Ann & Brown, Brian V., 2020, Forty-one new species of Coniceromyia Borgmeier (Diptera: Phoridae), an identification key, and new distributional records for the species of the genus, Zootaxa 4830 (1), pp. 1-61 : 20

Coniceromyia diadella sp. nov.

( Figs. 3B , 8N , 14B , 19C , 24O , 26F )

Diagnosis (male). Large wings (1.4 times longer than body length), wing thin veins and membrane deflected posteriorly ( Fig. 14B ), hind tibia apical quarter inflated with dorsal carina which continues along posteroapical margin, dense setulae posteriorly adjacent to the carina ( Fig. 26F ).

Material examined. Holotype ♂, ECUADOR: Napo: Volcán Sumaco , 0.55ºS, 77.68ºW, 14–16.iii.1996, yellow pan, 2100 m, P. Hibbs col. [LACM ENT 054937].

Description. Male. Body length, 2.5 mm. Head. Frons brown, shining, sparsely setulose, with weak median furrow. Flagellomere 1 light brown, conical very elongate. Arista could not be observed. Palpus light brown. Gena with two short setae; postocular seta short. Thorax. Scutum, scutellum, and pleural regions brown, shining; anepisternum bare. Legs narrow, elongated, yellowish-brown, except foretarsus and venter of forefemur brown. Foretibia with two small dorsal setae and anterodorsal row of strong setulae. Foremetatarsus with yellowish-brown anterior excavation, thick anteroapical setulose process, group of setae at excavation base, carina adjacent to excavation dorsal margin ( Fig. 8N ). Foremetatarsus ratio 0.3. Posterior face of hind femur with narrow basoventral band of tiny blunt setulae ( Fig. 19C ). Hind femur height/length ratio 0.23. Hind tibia with one dorsal seta in basal half, apical quarter inflated with dorsal carina which continues along posteroapical margin; dense setulae posteriorly adjacent to the carina. Wing ( Fig. 14B ). Large (1.4 times longer than body length), thin veins and membrane deflected posteriorly, membrane between apices of R 1 and R 4+5 darkened. Costa considerably swollen, 0.48 of the wing length; R 2+3 absent; M 1 slightly sinuous; M 2 nearly straight, CuA 1 -M 2 /M 2 -M 1 ratio 1.1; CuA 1 nearly straight. Halter white.

Abdomen. Tergites brown, with shining pubescence. Hypopygium light brown ( Fig. 24O ). Left epandrial process with a subepandrial wall extending from the hypoproct to its apex and a medially directed setulose projection near its base. Hypoproct with two setae.

Distribution. Known from a single mid-elevation site in Ecuador.

Etymology. Name derived from the Greek word diadelos (distinguishable, distinctive) and refers to the many uncommon features of this species such as its large wings and inflated hind tibia.

Figures

FIGURE 3. Habitus of the holotypes of new species of Coniceromyia, continuation (scale bars=1mm). A. C. deltopoda, B. C. diadela, C. C. diffusa, D. C. dolichopoda, E. C. hadrochaeta, F. C. hirtipenna.

FIGURE 8. Male foremetatarsus of new species of Coniceromyia (scale bars=0.1mm). A. C. amazonensis, B. C. anisocrossa, C. C. balbiae, D. C. bellatula, E. C. bisetosa, F. C. brachypoda, G. C. camptoneura, H. C. chrysopa, I. C. circulata, J. C. cladopyga, K. C. crassivena, L. C. dasypoda, M. C. deltopoda, N. C. diadela, O. C. diffusa, P. C. dolichopoda, Q. C. hadrochaeta, R. C. hirtipenna, S. C. inflata, T. C. leia, U. C. litopoda, V. C. luna, W. C. maculata, X. C. megalosoma, Y. C. memorialis, Z. C. niemeyeri, A’. C. numerosa, B’. C. ochyrosticha, C’. C. parallela, D’. C. pectinella.

FIGURE 14. Wings of the males of Coniceromyia new species, continuation. A. C. deltopoda, B. C. diadela, C. C. diffusa, D. C. dolichopoda, E. C. hadrochaeta, F. C. hirtipenna, G. C. inflata, H. C. leia, I. C. litopoda, J. C. luna, K. C. maculata, L. C. megalosoma.

FIGURE 19. Hind femora of the males of Coniceromyia new species, continuation (scale bars on figures of the entire femur=0.3 mm, on figures of ventrobasal part of the femur=0.1 mm). A–B. C. deltopoda, C. C. diadela, D–E. C. diffusa, F–G. C. dolichopoda, H–I. C. hadrochaeta, J. C. hirtipenna, K. C. inflata

FIGURE 24. Hypopygia of Coniceromyia new species in right lateral view (except figure J which is left lateral view). A. C. amazonensis, B. C. anisocrossa, C. C. balbiae, D. C. bellatula, E. C. bisetosa, F. C. brachypoda, G. C. camptoneura, H. C. chrysopa, I. C. circulata, J. C. cladopyga in left lateral view, K. C. cladopyga in right lateral view, L. C. crassivena, M. C. dasypoda, N. C. deltopoda, O. C. diadela, P. C. diffusa, Q. C. dolichopoda, R. C. hadrochaeta, S. C. hirtipenna, T. C. inflata, U. C. leia, V. C. litopoda, W. C. luna, X. C. maculata.

FIGURE 26. Unique or uncommon features of males of Coniceromyia species. A. Curved forecoxa of C. broadheadi, B–C, Sculpturing in dorsum of forefemur of C. broadheadi, D. Anterior view of foretibia of C. leia, showing the ventral face mostly bare and shiny but with differentiated thin setulae, E. Hind tibia of C. piricornis with dorsal patch of differentiated setulae, F. Hind tibia of C. diadella with dorsal carina with differentiated setulae, G. Posteroventral face of forefemur of C. systenopoda with a triangular ventral projection, H. Forefemur of C. pectinella with dorsal excavation bearing a comb of apically-directed curved setae, I. Foretrochanter of C. circulata with ventral process bearing apical group of curved setae, J. Foretrochanter of C. bellatula with ventral projection bearing apical group of curved setulae, K. Forefemur of C. chrysopa with bare oval ring surrounding an oval patch with small setulae, L. Forefemur of C. solaris with bare oval ring surrounding an oval patch with small setulae, M. Anterior face of midfemur of C. piricornis with ventral patch of dense setulae, N. Anterior face of midfemur of C. setosa with band of tiny setulae along basal two-thirds of ventral margin.