Coniceromyia luna, Ament & Kung & Brown, 2020

Ament, Danilo César, Kung, Giar-Ann & Brown, Brian V., 2020, Forty-one new species of Coniceromyia Borgmeier (Diptera: Phoridae), an identification key, and new distributional records for the species of the genus, Zootaxa 4830 (1), pp. 1-61 : 26-27

Coniceromyia luna sp. nov.

( Figs. 4D , 8V , 14J , 20 E–F, 24W, 27L)

Diagnosis (male). Flagellomere 1 globose, frons and thorax shining ( Fig. 4D ), foremetatarsus with bare, anterior excavation bearing small bumps, anterior preapical relatively long setulose process, several setulae at excavation base, and strong seta at base of anteroapical process ( Fig. 8V ).

Material examined. Holotype ♂, COLOMBIA: Boyacá: SFF Iguaque , 5.70ºN, 73.46ºW, 1–12.iv.2000, Malaise trap, 2975m, P. Reina col. ( IAVH) [LACM ENT 130609] . Paratypes: 3♂, same data as holotype ; 1♂, same but 17.iii–1.iv.2000; 1♂, Boyaca: SFF Iguaque: Cabaña Carrizal , 5.42ºN, 73.45ºW, 12.ix–13.x.2001 , Malaise trap, 2850m, P. Reina col.; 1♂, same but 30.vii–17.viii.2000; 1♂, same but 13.xi–4.xii.2000; 1♂, same but 10–28.vi.2001; 1♂, same but 4–21.xii.2000; 1♂, same but 7–24.ii.2001; 1♂, Cundinamarca: PNN Chingaza: Charrascales , 4.52ºN, 73.75ºW, 22.ii–15.iii.2002 , Malaise trap, 2990m, E. Raigoso col.; 2♂, Antioquia: San José de la Montaña: Vereda El Congo: Páramo El Congo, 4–15.xii.2016, Red pan trap, 3000-3200m, A. Sepulveda & C.H. Sepúlveda col. ( CEUA) ; 1♂, Antioquia: Belmira: Páramo de Sta Inés : Playitas Laguna, 06.62°N, 75.65°W, 4–14.xii.2016 , pan trap, 3200m, A.L. Montoya, J. Torres-Toro, J.P. Carmona col. ( CEUA) ; 2♂, same but El Morro , 6.63°N, 75.66°W, 2–12.xii.2017 , net, 2000-2500m, C. Rodríguez & A. Echeverry ( CEUA) ; 1♂, same but yellow pan trap.

Description. Male. Body length, 2.7 mm. Head. Frons brown, shining, with few sparse setulae; median furrow weak. Flagellomere 1 brown, globose. Arista subapical, about three times longer than flagellomere 1. Palpus brown to light brown. Gena with two to three setae. Thorax. Scutum, scutellum, and pleural regions brown to dark brown, shining; anepisternum bare. Scutum setae thin, posterior half rugose and almost bare. Foreleg yellowishbrown, except for brown foremetatarsus excavation, tarsomeres 4 and 5 and small rounded macula on dorsal apex of forefemur. Mid leg yellowish-brown, except apex of femur brown, base of tibia and tarsomeres 4 and 5. Hind leg light brown, except apical half of hind femur and hind tibia anteroapical region brown. Forefemur with posterobasal group of long setae. Foretibia with two dorsal setae and anterodorsal row of strong setulae. Foremetatarsus with bare, anterior excavation bearing small bumps, anterior preapical relatively long setulose process, several setulae at excavation base, and strong seta at base of anteroapical process ( Fig. 8V ). Foremetatarsus ratio 0.28. Hind femur expanded ventrally, with narrow basoventral band of relatively long setulae ( Fig. 20 E–F). Hind femur height/length ratio 0.46. Hind tibia with one dorsal seta in basal half. Wing ( Fig. 14J ). Dusky, covered with microtrichia longer than usual. Costa swollen, dark, 0.35 of the wing length; R 2+3 absent; M 1 slightly concave, slightly curved posteriorly at apex; M 2 slightly concave, CuA 1 -M 2 /M 2 -M 1 ratio 1.4; CuA 1 slightly sinuous. Halter white, brownish at base.

Abdomen. Tergites dark brown, with shining pubescence. Hypopygium brown ( Fig. 24W ), darkening surrounding epandrial setae sockets. Left epandrial process apically curved, with medially directed, curved, setose process near apex. Margin of right lobe of hypandrium without lateral projection. Hypoproct with two setae.

Distribution. High-elevation sites in Colombian Eastern and Western Andean Cordilleras ( Fig. 27L ).

Etymology. The specific epithet is Latin for moon and refers to the beautiful reflecting body of this species.

Variations. Specimens with a ventral row of slightly elongate setae on forefemur; specimens with a posterodorsal macula on forefemur.

Figures

FIGURE 4. Habitus of the holotypes of new species of Coniceromyia, continuation (scale bars=1mm). A. C. inflata, B. C. leia, C. C. litopoda, D. C. luna, E. C. maculata, F. C. megalosoma.

FIGURE 8. Male foremetatarsus of new species of Coniceromyia (scale bars=0.1mm). A. C. amazonensis, B. C. anisocrossa, C. C. balbiae, D. C. bellatula, E. C. bisetosa, F. C. brachypoda, G. C. camptoneura, H. C. chrysopa, I. C. circulata, J. C. cladopyga, K. C. crassivena, L. C. dasypoda, M. C. deltopoda, N. C. diadela, O. C. diffusa, P. C. dolichopoda, Q. C. hadrochaeta, R. C. hirtipenna, S. C. inflata, T. C. leia, U. C. litopoda, V. C. luna, W. C. maculata, X. C. megalosoma, Y. C. memorialis, Z. C. niemeyeri, A’. C. numerosa, B’. C. ochyrosticha, C’. C. parallela, D’. C. pectinella.

FIGURE 14. Wings of the males of Coniceromyia new species, continuation. A. C. deltopoda, B. C. diadela, C. C. diffusa, D. C. dolichopoda, E. C. hadrochaeta, F. C. hirtipenna, G. C. inflata, H. C. leia, I. C. litopoda, J. C. luna, K. C. maculata, L. C. megalosoma.

FIGURE 20. Hind femora of the males of Coniceromyia new species, continuation (scale bars on figures of the entire femur=0.3 mm, on figures of ventrobasal part of the femur=0.1 mm). A–B. C. leia, C–D. C. litopoda, E–F. C. luna, G–H. C. maculata, I. C. megalosoma, J. C. memorialis

FIGURE 24. Hypopygia of Coniceromyia new species in right lateral view (except figure J which is left lateral view). A. C. amazonensis, B. C. anisocrossa, C. C. balbiae, D. C. bellatula, E. C. bisetosa, F. C. brachypoda, G. C. camptoneura, H. C. chrysopa, I. C. circulata, J. C. cladopyga in left lateral view, K. C. cladopyga in right lateral view, L. C. crassivena, M. C. dasypoda, N. C. deltopoda, O. C. diadela, P. C. diffusa, Q. C. dolichopoda, R. C. hadrochaeta, S. C. hirtipenna, T. C. inflata, U. C. leia, V. C. litopoda, W. C. luna, X. C. maculata.

FIGURE 27. Distributional records of Coniceromyia new species (species known from a single or close localities were not illustrated. Squares=holotype, circles=other specimens). A. C. amazonensis, B. C. bellatula, C. C. bisetosa, D. C. brachypoda, E. C. cladopyga, F. C. dasypoda, G. C. deltopoda, H. C. diffusa, I. C. hadrochaeta, J. C. hirtipenna, K. C. litopoda, L. C. luna, M. C. maculata.