Coniceromyia megalosoma, Ament & Kung & Brown, 2020

Ament, Danilo César, Kung, Giar-Ann & Brown, Brian V., 2020, Forty-one new species of Coniceromyia Borgmeier (Diptera: Phoridae), an identification key, and new distributional records for the species of the genus, Zootaxa 4830 (1), pp. 1-61 : 28

Coniceromyia megalosoma sp. nov.

( Figs. 4F , 8X , 14L , 20I , 25A , 28A )

Diagnosis (male). Large (more than 3.5 mm of body length), legs narrow, elongated, hind femur yellowish-brown, except dorsal and ventral margin dark brown and apex brown ( Fig. 4F ), hypopygium left epandrial process apically bifurcated into a dorsal and a ventral lobe.

Material examined. Holotype ♂, ECUADOR: Napo: Oyacachi , 0.22ºS, 78.08ºW, 19.ii–29.iii.1996, Malaise trap, 3000 m, P. Hibbs col. [LACM ENT 021055] . Paratypes: COLOMBIA: 2♂, Antioquia: Belmira: Paramo de Sta. Inés, El Morro , 06.63°N, 75.64°W, 2–12.xii.2017, Malaise trap dossel, 3100–3300m, C Rodríguez & A. Echeverry ( CEUA) ; 7♂, same but 25.v–5.iv.2017, A.L. Montoya, C. Rodriguez & J. P. Carmona ( CEUA) ; 5♂, same but Playitas Laguna , 6.64°N, 75.67°W, 4–14.xii.2016, Malaise trap, 2500–3000m, A.L. Montoya, J. Torres-Toro, J.P. Carmona col. ( CEUA) .

Description. Male. Body length, 3.7 mm. Head. Frons dark brown, shining, sparsely setulose, with well-defined median furrow. Flagellomere 1 brown, conical elongate. Arista apical, subequal length of flagellomere 1. Palpus light brown. Gena with two setae. Thorax. Scutum, scutellum, and pleural regions brown, shining; scutum posterior half rugose; anepisternum bare. Legs narrow, elongated. Fore and mid legs yellowish-brown, except brown anteroventral maculae on fore and mid femur and on fore and mid coxae. Hind femur yellowish-brown, except dorsal and ventral margin dark brown and apex brown; hind tibia and tarsus brown. Foretibia with two dorsal setae and anterodorsal row of strong setulae. Foremetatarsus with anterior excavation bearing row of short strong setulae, anteroapical setulose process, group of large setae at base of excavation, and strong seta at base of anteroapical process ( Fig. 8X ). Foremetatarsus ratio 0.18. Posterior face of hind femur with elongate basoventral band of tiny blunt setulae, broad basally tapering toward and along ventral margin, extending two-thirds of the hind femur ( Fig. 20I ). Hind femur height/length ratio 0.28. Hind tibia with one dorsal seta in basal half. Wing ( Fig. 14L ). Dusky. Costa and radial veins considerably swollen, Costa 0.45 of the wing length; R 2+3 absent; M 1 slightly concave, curved posteriorly at apex; M 2 sinuous, CuA 1 -M 2 /M 2 -M 1 ratio 2.1; CuA 1 slightly sinuous. Halter base brown, apex light brown. Abdomen. Tergites brown, with shining pubescence. Hypopygium brown ( Fig. 25A ). Left epandrial process apically bifurcated into a dorsal and a ventral lobe, with a subepandrial elevation bearing apical setae. Right subepandrial plate developed posteriorly, curved ventrally. Hypoproct without setae.

Distribution. High-elevation sites in Equatorian and Colombian Andes ( Fig. 28A ).

Etymology. Name derived from the Greek roots megale (large) and soma (body).

Figures

FIGURE 4. Habitus of the holotypes of new species of Coniceromyia, continuation (scale bars=1mm). A. C. inflata, B. C. leia, C. C. litopoda, D. C. luna, E. C. maculata, F. C. megalosoma.

FIGURE 8. Male foremetatarsus of new species of Coniceromyia (scale bars=0.1mm). A. C. amazonensis, B. C. anisocrossa, C. C. balbiae, D. C. bellatula, E. C. bisetosa, F. C. brachypoda, G. C. camptoneura, H. C. chrysopa, I. C. circulata, J. C. cladopyga, K. C. crassivena, L. C. dasypoda, M. C. deltopoda, N. C. diadela, O. C. diffusa, P. C. dolichopoda, Q. C. hadrochaeta, R. C. hirtipenna, S. C. inflata, T. C. leia, U. C. litopoda, V. C. luna, W. C. maculata, X. C. megalosoma, Y. C. memorialis, Z. C. niemeyeri, A’. C. numerosa, B’. C. ochyrosticha, C’. C. parallela, D’. C. pectinella.

FIGURE 14. Wings of the males of Coniceromyia new species, continuation. A. C. deltopoda, B. C. diadela, C. C. diffusa, D. C. dolichopoda, E. C. hadrochaeta, F. C. hirtipenna, G. C. inflata, H. C. leia, I. C. litopoda, J. C. luna, K. C. maculata, L. C. megalosoma.

FIGURE 20. Hind femora of the males of Coniceromyia new species, continuation (scale bars on figures of the entire femur=0.3 mm, on figures of ventrobasal part of the femur=0.1 mm). A–B. C. leia, C–D. C. litopoda, E–F. C. luna, G–H. C. maculata, I. C. megalosoma, J. C. memorialis

FIGURE 25. Hypopygia of Coniceromyia new species in right lateral view, continuation. A. C. megalosoma, B. C. memorialis, C. C. niemeyeri, D. C. numerosa, E. C. ochyrosticha, F. C. parallela, G. C. pectinella, H. C. platypoda, I. C. polychaeta, J. C. puntarenensis, K. C. riccardiae, L. C. setosa, M. C. solaris, N. C. stichochaeta, O. C. strongyla, P. C. systenopoda, Q. C. trilobata, R. C. xiei.

FIGURE 28. Distributional records of Coniceromyia new and previously known species, continuation (species known from a single or close localities were not illustrated). Squares=holotype, circles=other specimens). A. C. megalosoma, B. C. niemeyeri, C. C. numerosa, D. C. parallela, E. C. puntarenensis, F. C. stichochaeta, G. C. trilobata, H. C. affinis, I. C. anacleti, J. C. apicalis, K. C. apioneura, L. C. auristriata, M. C. basalis, N. C. brevivena, O. C. broadheadi, P. C. browni, Q. C. convergens, R. C. costaricana, S. C. disparivena, T. C. epicantha, U. C.femoralis, V. C. franciscana.