Atanatolica angulata , Razuri-Gonzales, Ernesto, Holzenthal, Ralph W & Rios-Touma, Blanca, 2018

Razuri-Gonzales, Ernesto, Holzenthal, Ralph W & Rios-Touma, Blanca, 2018, New Atanatolica species from Ecuador (Trichoptera, Leptoceridae), ZooKeys 793, pp. 97-114: 97

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Atanatolica angulata

sp. n.

Atanatolica angulata  sp. n. Figs 3, 8


This species is related to A. aurea  Holzenthal, 1988 from Colombia, and A. penai  Holzenthal, 1988 from Bolivia, especially in the broad apicomesal processes of tergum X. The new species differs from A. aurea  by the much shorter, acute, angulate apicomesal processes, and the rounded apicolateral processes on tergum X as well as the shape of the inferior appendages, especially the posteromesal margin of the first segment of this structure in ventral view. From A. penai  , it differs by the shape of the apicomesal processes of tergum X, which in A. angulata  sp. n. are strongly angulate. They also differ by the shape of the inferior appendages, which have a mesal bump on the inner surface in A. penai  , but not in A. angulata  sp. n.


Adult male. Forewing length 9 mm (n = 1). General color light brown, forewing membrane brown, covered in golden and brown setae. Head with long, white setae. Antennae with long, white setae on scape and pedicel, flagellomeres with dark brown setae and ring of white setae basally. Maxillary palps light brown, with long, white and short brown hairs. Thorax light brown with brown hairs. Forelegs with coxae, trochanter, and femur light brown with long, white setae; tibia and tarsomeres with white and brown setae; mid and hind legs light brown with white and brown setae, tibia and tarsomeres with two rows of dark spines ventrally. Tibial spur formula 0, 2, 2.


Segment IX annular, short, with anterior and posterior margins sinuous, setae on ventral and lateral surfaces not associated with warts (Figure 3A). Preanal appendages slightly shorter than tergum X, slender, digitate, setose (Figure 3A, 3B). Tergum X notched apicomesally, basal portion of tergum X membranous, with two pairs of sclerotized apical processes; apicomesal processes flattened, directed dorsad in lateral view (Figure 3A), strongly angulate, posterolateral apex acute in dorsal and caudoventral views (Figure 3B,) ; apicolateral processes capitate, directed dorsad in lateral view, slightly longer than the apicomesal processes, but equally as long in dorsal and caudoventral views (Figs 3A, B, D), digitate in dorsal and caudoventral views (Figure 3B, D). Inferior appendages with basal portion of first segment broad, setose, mesal margin forming an angle, apical portion elongate, digitate, slightly curved, and slightly inflated basally in ventral view (Figure 3C), covered with stout, spine-like setae on mesal surface; second article apparently fused with the first segment, represented by triangular semimembranous process (Figure 3A, C). Phallic apparatus simple, without any processes; phallobase short, tubular, curved downwards; phallotremal sclerite simple, spine-like, curved; endothecal membranes not everted (Figure 3E).

Holotype male.

ECUADOR: Napo: Río Jondachi, 30 km N Tena, 950 m, 10.ix.1990, OS Flint ( NMNH) [USNMENT01295341].


The specific name angulata  is a Latin adjective referring to the angulate apicomesal processes on tergum X.


Napo Province (Ecuador) (Figure 8).