Heptathela tokashiki, Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang & Li, Daiqin, 2019

Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang & Li, Daiqin, 2019, A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands, ZooKeys 888, pp. 1-50 : 1

Heptathela tokashiki sp. nov. Fig. 21

Type material.

Holotype: JAPAN · ♀; Okinawa-ken, Tokashikijima Island, Aharen; 26.19N, 127.37E; alt. 100 m; 8 May 2014; D. Li and B. Wu leg.; XUX-2014-062.

Paratypes: JAPAN · 25 ♀♀; same data as for holotype; XUX-2014-046 to 063D · 3 ♀♀; same data as for holotype; 27 December 2012; D. Li, F.X. Liu and X. Xu leg.; XUX-2012-417, 421, 425.

Diagnosis.

Females of H. tokashiki sp. nov. can be distinguished from H. shuri sp. nov. by the long inner receptacular clusters with several granules ( Fig. 21 A–D ). Heptathela tokashiki sp. nov. can also be diagnosed from all other Okinawa group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: T (68), A (200), C (290), A (362).

Description.

Female (Holotype). Carapace brown; opisthosoma dark brown, with dark brown tergites close to each other; cheliceral groove with eleven pronounced denticles; seven spinnerets. Measurements: BL 11.80, CL 5.15, CW 4.10, OL 6.80, OW 5.10; ALE> PLE> PME> AME; palp 8.60 (3.05 + 1.50 + 1.95 + 2.10), leg I 10.50 (3.35 + 1.85 + 2.15 + 2.10 + 1.05), leg II 10.45 (3.40 + 1.75 + 2.00 + 2.20 + 1.10), leg III 10.95 (3.20 + 1.85 + 1.80 + 2.60 + 1.50), leg IV 15.70 (4.60 + 2.00 + 2.80 + 3.90 + 2.40).

Female genitalia. A pair of depressions on the ventro-lateral part of the genital atrium ( Fig. 21B, D ). Paired receptacular clusters along the anterior margin of bursa copulatrix, divided into two parts, the inners longer than the laterals, both with several granules, without genital stalks ( Fig. 21A, B ).

Male. Unknown.

Etymology.

The species epithet, a noun in apposition, refers to the type locality.

Distribution.

The species is endemic to the Japanese island Tokashikijima ( Fig. 1C ).

Figures

Figure 1. General somatic morphology of Heptathela kimurai (Kishida, 1920) A dorsal view B ventral view and map showing the sampling localities of Heptathela specimens from Kyushu to central Ryukyus (C).

Figure 21. Female genital anatomy of Heptathela tokashiki sp. nov. A, B 4062 (holotype, short for XUX- 2014 - 062) C, D 4063 A, C vulva dorsal view B, D vulva ventral view. Scale bar: 0.5 mm.