Archaeoripiphorus nuwa Hsiao, Yu & Deng

Hsiao, Yun, Yu, Yali, Deng, Congshuang & 4, Hong Pang, 2017, The first fossil wedge-shaped beetle (Coleoptera, Ripiphoridae) from the middle Jurassic of China, European Journal of Taxonomy 277, pp. 1-13: 4-8

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scientific name

Archaeoripiphorus nuwa Hsiao, Yu & Deng

sp. nov.

Archaeoripiphorus nuwa Hsiao, Yu & Deng   sp. nov.

Figs 1–4 View Fig View Fig View Fig View Fig


As for the genus (vide supra). In particular this species resembles the extant ptilophorine species, Trigonodera tokejii (Nomura & Nakane, 1959)   in general appearance ( Fig. 5 View Fig ), from which it differs by

its shorter antennae and terminal maxillary palpomere, wider pronotum anteriorly, shorter elytra, and tibial edges without spiniform seta apically.


The specific name honors the great goddess in the ancient Chinese mythology, Nüwa, who created mankind and saved humanity from a major catastrophe by using the five-colored stones to repair the damage to the pillar of heaven caused by the water god, Gonggong.

Type material


CHINA: nearly complete specimen in dorsal position; sex unknown   ( CNU-C-NN-2006841).


Type locality

CHINA: NE China, Daohugou, Shantou Township, Ningcheng County, Inner Mongolia.

Type horizon

Jiulongshan Formation, Middle Jurassic, about 165 Ma ( Gao & Ren 2006).


BODY ( Figs 1–2 View Fig View Fig ). Length 15.5 mm, width 5.0 mm, head length 2.0 mm, antennal length 3.4 mm, pronotum length 3.5 mm, pronotum width 4.0 mm, elytra length 10.0 mm. Surface densely covered with fine pubescence ( Fig. 3E View Fig ).

HEAD ( Fig. 3A View Fig ). Small and elongate, abruptly constricted posteriorly to form broad neck; surface sparsely punctate. Eyes oval, shallowly emarginate, distinctly separated from each other, ratio of eye diameter to interocular space 1.00:1.65; surface finely faceted. Frontoclypeal suture indistinct. Apical margin of clypeus slightly sinuate. Labrum rounded, subquadrate. Terminal maxillary palpomere elongate securiform, about 4.0 times as long as minimum width. Antennal insertions widely separated, exposed from above. Antennae 11-segmented. Scape and pedicel very short, antennomere III strongly elongate, IV to X rectangular or trapezoidal, antennomere XI with pointed apex ( Fig. 3D View Fig ). Length ratio of antennomeres as follows: 1.20:1.00:2.30:1.35:1.20:1.35:1.10:0.90:0.90:0.90:1.45.

PROTHORAX ( Fig. 3B View Fig ). Almost triangular, about 0.88 times as long as wide, slightly narrower than elytra at base, tapering anteriorly; anterior margin truncate and straight; anterior angles rounded and indistinct, posterior margin trilobed, with posterior angles moderately protruding and obtuse; dorsal surface densely punctate. Scutellar shield minute, angulate apically.

ELYTRA ( Fig. 3C View Fig ). About 2 times as long as wide, lateral sides slightly narrowing posteriorly; surface densely and regularly punctate. Abdomen with five tergites and ventrites.

LEGS. Long and slender; tibiae moderately widening apically, without spiniform seta at apices, tibial spur formula 1-1-2 ( Fig. 4 View Fig A–F); tarsi slender, tarsal formula 5-5-4; tarsus nearly as long as tibiae in fore and mid legs, slightly shorter than tibiae in hind legs; tarsomeres cylindrical, hind tarsomere I slightly longer than tarsomeres II and III combined. Claws slightly pectinate ( Figs 4 View Fig G–H).