Calendula suffruticosa subsp. riffensis A.C. Gonç. & P. Silveira, 2023
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publication ID |
https://doi.org/ 10.11646/phytotaxa.605.1.1 |
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persistent identifier |
https://treatment.plazi.org/id/E82AFC0A-157D-CA5B-66ED-FB10855AFD0D |
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treatment provided by |
Plazi |
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scientific name |
Calendula suffruticosa subsp. riffensis A.C. Gonç. & P. Silveira |
| status |
subsp. nov. |
11k. Calendula suffruticosa subsp. riffensis A.C. Gonç. & P. Silveira , subsp . nov. Type:— MOROCCO. Tetouan , Jbel
Kelti, Arifane, 943 m, [35°17’04” N, 5°18’00” W], 15 May 2014, P. Silveira & A. C. R. S. Gonçalves 3341 (holotype GoogleMaps
RAB!; isotypes AVE!, MARK!).
Diagnosis:— Calendula suffruticosa subsp. riffensis differs from C. suffruticosa subsp. suffruticosa by the young parts ± white-arachnoid, as opposed to being entirely ± glabrous to glandulose, and by the cymbiform aquenes width (5,0) 5,2–5,5 (5,8) mm, as opposed to (6.7) 7.7–10.8 (11.8) mm. It differs from subsp. boccoyana by having basal leaves with a ratio length/width of (2.2) 2.4–4.0 (4.4) instead of (4.9) 5.0–6.3 (6.5) and a thickness of (0.2) 0.3–0.4 (0.6) mm, instead of (0.3) 0.4–0.6 (0.8) mm.
Description:––Perennial herbs. Stems (28) 32–49 (71) cm long, erect to ascending with glandular hairs distally and predominance of non-glandular white-arachnoid pubescence proximally and in young branches. Basal leaves (4.1) 4.5–8.2 (8.3) × 1.5–2.1 (2.3) cm, (0.2) 0.3–0.4 (0.6) mm thick, obovate to oblong, apex obtuse, margins sub-entire to slightly sinuate-dentate, lamina with more glandular than non-glandular hairs, except in margins or leaves of young branches where non-glandular white-arachnoid pubescence predominates. Capitula solitary, (2.9) 3.2–3.6 (4) cm diameter. Outer achenes rostrate 17–21 (22) × (1.2) 1.3–1.5 (2.8) mm, ± straight or slightly curved, without dorsal spines, without a ventral basal tooth; middle achenes bialate (10) 10.8–15 × (5.8) 6.7–10 mm, with a rostrum (5) 5.4–7.9 (8.7) mm, margin of lateral wings sinuate-dentate to incised; cymbiform (5,8) 7,5–10,5 (11,3) × (5,0) 5,2–5,5 (5,8) mm; inner achenes vermiculate-alate (4,3) 4,5–5,8 (6,3) × (2,5) 2,8–4,5 (5) mm; and vermiculate-exalate 3,3–4,7 (5) × 1,5–1,7 (1,8) mm, both circular. Figure 17 View FIGURE 17 .
Habitat and distribution:—Limestone rocks/cliffs, at elevations from 30 to 1800 m. It is found in perhumid Mediterranean bio-climate. Endemic to Morocco with a restricted distribution in the southern Tetouan Mountains (Arrifane – Jbel Kelti).
Conservation status:— Data Deficient (DD) . This taxon is under numerous threats, especially due to climate change (extreme weather), and droughts. The taxon is rare and local with a small-restricted range; only one population is known, and few herbarium specimens exist (from the same locality). The number of mature individuals in this population is estimated to be <250 based on field observations. The AOO is 4 km 2. However, there is inadequate information to assess their extinction risk based on distribution, since it is possible that other subpopulations exist in the area.
Chromosome number:—2n = 32.
Genome size:—3.06 ± 0.10 pg.
Eponymy:—For the Riff region in northern Morocco where it occurs.
Additional collections:— MOROCCO. Tanger-Tetouan-Al Hoceïma: Chefchaouene c. 1 km SW of Oued Laou (town), on main road, 30 m, [35°26’14” N, 5°04’54” W], 26 February 1994, S. L GoogleMaps . Jury, A . Taleb, T. M . Upson & G. S . Walters 13384 ( BM!); Chefchaouene Chefchauen , Targa, Oued Tarerha, montes Beni-Galoune, [35°23’42” N, 5°00’37” W], 4 May 1996, M. A GoogleMaps . Mateos, F. J . Pina & S. Silvestre MM 422 96 ( BC!); Chefchaouene track N from Chefchauen around Jbel El Kelaa at village Ighrmane, 720 m, [35°14’09” N, 5°13’17” W], 8 June 2005, M. A GoogleMaps . Lafkih, M . Carine, S. L . Jury & F. J . Rumsey 244 ( BM!); Chefchaouene Tetouan , c. 38 km from Chefchaouen, 14 km above Bab Taza on track to Jbel Talamssmtane, 1765 m, [35°01’54” N, 5°16’00” W], 26 June 1992, Optima Inter V GoogleMaps 2173 ( RNG, PAL!); Chefchaouene , carretera Oued Laou, 220 m, [35°10’08” N, 5°15’49” W], 15 May 1990, B GoogleMaps . Cabezudo & J. M . Nieto s.n. ( MGC!) .
| P |
Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants |
| A |
Harvard University - Arnold Arboretum |
| C |
University of Copenhagen |
| R |
Departamento de Geologia, Universidad de Chile |
| S |
Department of Botany, Swedish Museum of Natural History |
| L |
Nationaal Herbarium Nederland, Leiden University branch |
| T |
Tavera, Department of Geology and Geophysics |
| M |
Botanische Staatssammlung München |
| G |
Conservatoire et Jardin botaniques de la Ville de Genève |
| BM |
Bristol Museum |
| F |
Field Museum of Natural History, Botany Department |
| J |
University of the Witwatersrand |
| BC |
Institut Botànic de Barcelona |
| N |
Nanjing University |
| V |
Royal British Columbia Museum - Herbarium |
| RNG |
University of Reading, Centre for Plant Diversity and Systematics |
| PAL |
Herbarium Mediterraneum Panormitanum |
| B |
Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet |
| MGC |
Universidad de Málaga |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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