Amberana Distant, 1908

Bucher, Manon, Bouteille, Emilien & Cesne, Maxime Le, 2024, Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar, European Journal of Taxonomy 935, pp. 166-202 : 174-175

Key to species of Amberana Distant, 1908

1. Tegmina mainly red, orange or coral ................................................................................................ 2

– Tegmina other than with a red or orange dominant colour ............................................................... 9

2. Thorax with uniform colour, red or black ......................................................................................... 3

– Thorax with more than one colour .................................................................................................... 4

3. Thorax black, total length less than 11.0 mm ................................................................................... 5

– Thorax red, total length longer than 11.0 mm .................................................................................. 7

4. Thorax black with transversal orange band on pronotum ........... A. boucomonti ( Lallemand, 1920)

– Thorax coloured coral with apex of scutellum black .......................................................................... ................................................................. A. harandranomasina Le Cesne & Soulier-Perkins, 2022

5. Total length not exceeding 8.0 mm ................................................................................................... 6

– Total length longer than 9.0 mm ............................ A. peyrierasi Soulier-Perkins & Le Cesne, 2022

6. Black border following costal, apical and sutural margins of tegmina, from tip of clavus black border get narrower and follows shortly clavo-corial suture. Sometimes, a black patch can extend from the tip of clavus down to the costal margin ................................................. A. limbata Lallemand, 1955

– Black border on tegmina, starting only from tip of clavus, following apical margin and stopping at mid length of costal margin ..................................................................... A. pygmaea Schmidt, 1920

7. Black border starting from tip of clavus and following apical and costal margins of tegmina .......... ................................................................................................................ A. marginata ( Fallou, 1890)

– Black line running along clavo-corial suture ................................................................................... 8

8. Black border all around tegmina, thin black line on clavo-corial suture and a black line on corium starting after M + Cu split and finishing before reaching apex .................. A. uncinata Jacobi, 1917

– Larger black border around tegmina but finishing before last third of costal margin, black line on clavo-corial suture large, line on corium not individualized ...................... A. fissurata Jacobi, 1917

9. Tegmina partially translucent or smoked ........................................................................................ 10

– Tegmina opaque .............................................................................................................................. 13

10. Tegmina without a black border following its apical margin .............. A. dimidiata ( Signoret, 1860)

– Black border following apical margin of tegmina ...........................................................................11

11. Tegmina translucent with an orange patch along clavus basal margin and red area along base of costal margin .................................................................................. A. noualhieri ( Lallemand, 1920)

– Tegmina yellowish smoked ............................................................................................................ 12

12. Red patch at base of tegmina plus red spot within internal curve of black apical border; vertex and pronotum red ............................................................................. A. maromandiana Lallemand, 1949

– Orange patch at base of tegmina bordered posteriorly by perpendicular black line that follows clavo-corial suture within orange patch; vertex and scutellum black, pronotum orange ............................. ................................................................................................................... A. lemuria ( Distant, 1908)

13. Pronotum completely black ............................................................................................................ 14

– Pronotum black bearing a yellow to orange transversal band ........................................................ 21

14. Tegmina mainly cream, with a black border and some red .......... A. ouvrardi Soulier-Perkins, 2012

– Tegmina mainly dark brown to black ............................................................................................. 15

15. Tegmina with anterior third orange and rest black ................................ A. humeralis ( Fallou, 1890)

– Tegmina dark brown to black, bearing 2 or 3 spots of colour ........................................................ 16

16. Total length not exceeding 7.0 mm ................................................................................................. 17

– Total length longer than 7.0 mm ..................................................................................................... 18

17. Tegmina bearing three yellow spots, the two on corium are small ..... A. attiei Soulier-Perkins, 2012

– Tegmina bearing two patches of colour orange ........................................... A. kraussi Synave, 1957

18. Total length inferior to 11.0 mm ..................................................................................................... 19

– Total length equal or longer than 11.98 mm ................................... A. pascali Soulier-Perkins, 2012

19. Apex of subgenital plate pointed ..................................................... A. bergevini ( Lallemand, 1920)

– Apex of subgenital plate slightly bulbous ................................................. A. elongata Distant, 1908

20. Tegmina bearing only band of colour along anterior claval margin, rest of tegmina dark brown with some red ........................................................................................ A. brolemanni ( Lallemand, 1920)

– Tegmina bearing band of colour along anterior claval margin plus two other spots of same colour ..... ......................................................................................................................................................... 21

21. Clypeus black .................................................................................................................................. 22

– Clypeus yellow to orange .................................................................. A. sexguttata (Melichar, 1915)

22. Tegmina with an orange spot on the clavus and two on the corium ....... A. melanops ( Jacobi, 1917)

– Tegmina with two longitudinal yellow lines, one on clavus and one on corium ................................ ......................................................................................... A. vittipennis ( Bergroth, 1894) comb. nov.