Draconarius Ovtchinnikov, 1999: 70

WANG, XIN-PING, 2002, A Generic-Level Revision Of The Spider Subfamily Coelotinae (Araneae, Amaurobiidae), Bulletin of the American Museum of Natural History 2002 (269), pp. 1-150 : 66-73

Draconarius Ovtchinnikov, 1999: 70 .

TYPE SPECIES: Draconarius venustus Ovtchinnikov, 1999 , by original designation.

DIAGNOSIS: Resembles Asiacoelotes in having an elongated cymbial furrow, a long, slender embolus, and long, strongly convoluted spermathecae. Females can be distinguished by the posteriorly originated copulatory ducts and the widely separated spermathecae (figs. 182, 193); males by having a conductor dorsal apophysis (figs. 184, 185, 198).

DESCRIPTION: See description of Draconarius wudangensis (Chen and Zhao) .

DISTRIBUTION: Tadzhikistan, Bhutan, Nepal, China, Korea (map 7).

COMPOSITION: Twenty­six species are included. All are new combinations except the type species.

The species Draconarius wudangensis ( Chen and Zhao, 1997) and Draconarius sinualis ( Chen, Zhao and Wang, 1991) are described here. The male of D. sinualis is described for the first time.

1. Draconarius arcuatus ( Chen, 1984) : female paratypes from Huanglongdong, Hangzhou, Zhejiang, China, in HTC, examined. NEW COMBINATION (transferred here from Coelotes ).

2. Draconarius aspinatus ( Wang et al., 1990) : female holotype, male and female paratypes from Huangshan, Anhui, China, in HBI, examined. NEW COMBINATION (transferred here from Coelotes ).

3. Draconarius baccatus ( Wang, 1994) : male and female paratypes from Xiangfan, Hubei, China, in HBI, examined. NEW COM­ BINATION (transferred here from Coelotes ).

4. Draconariuss baronii (Brignoli, 1978) : male holotype only, female paratype is a different species (of the genus Himalcoelotes ). Male holotype from Dorjula, Bhutan, in NHMB, examined. NEW COMBINATION (transferred here from Coelotes ).

5. Draconarius calcariformis ( Wang, 1994) : female holotype and male paratype form Dabashan, Hubei, China, in HBI, examined. NEW COMBINATION (transferred here from Coelotes ).

6. Draconarius cheni ( Platnick, 1989) : female holotype and paratype from Huanglongdong, Hangzhou, Zhejiang, China, in HTC, examined. NEW COMBINATION (transferred here from Coelotes ).

7. Draconarius coreanus ( Paik and Yaginuma, 1969) : female holotype, male and female paratypes from Seong­gul Cave, Korea, in NSMT, not examined. NEW COMBINATION (transferred here from the genus Coelotes ). Male and female specimens from Korea, in KAI, examined.

8. Draconarius davidi ( Schenkel, 1963) : female holotype from Inkiaphou, Shensi, China, in MNHN, examined. NEW COMBINA­ TION (transferred here from Coelotes ).

9. Draconarius gurkha ( Brignoli, 1976) : female holotype from Yak­Alm, Taboche, Nepal, in ZII, examined. NEW COMBINATION (transferred here from Coelotes ).

The species Coelotes lama Brignoli, 1976 : female holotype from Nepal, in ZII, examined, has the identical genitalic morphology and is placed as a junior synonym of D. gurkha (Brignoli) . NEW SYNONYMY (transferred here from Coelotes ).

10. Draconarius infulatus ( Wang et al., 1990) : female holotype and 1 female paratype from Tianmushan, Zhejiang, China, in HBI, not examined. NEW COMBINATION (transferred here from Coelotes ).

11. Draconarius labiatus ( Wang and Ono, 1998) : female holotype and paratype from Nantou, Taiwan, in NSMT, examined. NEW COMBINATION (transferred here from Coelotes ).

12. Draconarius lutulentus ( Wang et al., 1990) : male and female paratypes from Zhangjiajie, Dayong, Hunan, China, in HBI, examined. NEW COMBINATION (transferred here from Coelotes ).

13. Draconarius molluscus ( Wang et al., 1990) : female paratypes from Lushan, Ji­ angxi, China, in HBI, examined. NEW COM­ BINATION (transferred here from Coelotes ).

14. Draconarius neixiangensis (Hu, Wang and Wang, 1991) : female holotype and male and female paratypes from Neixiang, Henan, China, deposited in Shandong University, Ji­

nan, China, not examined. NEW COMBINATION (transferred here from Coelotes ).

15. Draconarius potanini ( Schenkel, 1963) : female holotype from Gansu, China, in MNHN, examined. NEW COMBINATION (transferred here from Cybanus).

16. Draconarius schenkeli (Brignoli, 1978) : female holotype from Chimakothi, Bhutan, in NHMB, examined. NEW COMBI­ NATION (transferred here from Coelotes ).

17. Draconarius singulatus ( Wang et al., 1990) : male and female paratypes from Nan­ shanping, Chenbu, Hunan, China, in HBI, examined. NEW COMBINATION (transferred here from Coelotes ).

18. Draconarius sinualis ( Chen, Zhao and Wang, 1991) : female holotype from Jinding, Wudangshan, Hubei, China, in HUW, examined. NEW COMBINATION (transferred here from Coelotes ). Both male and female specimens from China (Hubei), in IZB, examined.

19. Draconarius stemmleri (Brignoli, 1978) : female holotype and paratype from Sha Gogona, Bhutan, in NHMB and MCV, examined. NEW COMBINATION (transferred here from Coelotes ).

20. Draconarius striolatus ( Wang et al., 1990) : female holotype from Yuzhong, Gansu, China, in HBI, examined. NEW COMBINA­ TION (transferred here from Coelotes ).

21. Draconarius trifasciatus ( Wang and Zhu, 1991) : female holotype and male paratype from Mt. Emei, Sichuan, China, in NBUMS, not examined. NEW COMBINATION (transferred here from Coelotes ).

22. Draconarius venustus Ovtchinnikov, 1999 : male holotype and female paratype from Yachsu River valley, Khazratisho Mt., Tadzhikistan, in CSO, not examined.

23. Draconarius wenzhouensis ( Chen, 1984) : female holotype and paratype from Xueshan, Wenzhou, Zhejiang, China, in HTC, examined. NEW COMBINATION (transferred here from Coelotes ).

24. Draconarius wudangensis ( Chen and Zhao, 1997) : male and female paratypes from Jinding, Wudangshan, Hubei, China, in HUW, examined. NEW COMBINATION (transferred here from Coelotes ).

25. Draconarius wuermlii (Brignoli, 1978) : female holotype from Dechhi Paka, Bhutan, in NHMB, examined. NEW COMBI­ NATION (transferred here from Paracoelotes ).

26. Draconarius yosiianus (Nishikawa, 1999) : female holotype and paratype from Jiabao Dong, Xingren Cun, Jiazhuan Xiang, Bama Xian, northwestern Guangxi, China, in NSMT, not examined. NEW COMBINATION (transferred here from Coelotes ).

Draconarius wudangensis (Chen and Zhao) Figures 181–191

Coelotes wudangensis Chen and Zhao, 1997: 87 , figs. 1–4 (male and female paratypes from Jind­ ing, Wudangshan, Hubei, China, in HUW, ex­

amined). – Song et al., 1999: 388, figs. 226K,

226L, 227O, 229A.

DIAGNOSIS: This species is similar to D. venustus but can be distinguished by the large, anteriorly situated spermathecal heads of females (fig. 182), the relatively short RTA, the less extended cymbial furrow, and the different shape of embolic base of males (figs. 183–185).

DESCRIPTION: Total length 6.00 –12.0. From front, anterior eye row slightly procurved, posterior row procurved; eye sizes and arrangements: AME smallest, ALE largest, PLE slightly larger than PME; AME­AME slightly shorter than their diameter, AME­ ALE separated by roughly AME radius, ALE­PLE close together, PME­PLE separat­ ed by more than their diameter, PME­PME slightly wider than their diameter. Clypeal height about twice AME diameter or slightly less; chilum divided, elongated. Chelicerae with three promarginal and two retromarginal teeth. Labium slightly shorter than wide. Length of female 1st leg patella + tibia longer than carapace length.

Tarsal organ situated dorsally, away from distal end of tarsus, close to second or third distal trichobothrium (figs. 186–188). Female spinnerets examined: apex of ALS with 2 major ampullate gland spigots (MAP), about 46 piriform gland spigots; PMS not well prepared, with 1 minor ampullate gland spigots visible, at least 20 aciniform gland spigots, and 1 cylindrical gland spigot (CY) observed; PLS with about 34 aciniform gland spigots and 2 cylindrical gland spigots (figs. 189–191).

Epigynum with epigynal teeth short, widely separated, situated anterior of atrium; atrium small; copulatory ducts originated posteriorly, mesal of spermathecae, more or less elongated; spermathecal heads apparent; spermathecae broad, elongated, convoluted, with indistinct stalks and bases (figs. 181, 182). Male palp with patellar apophysis long; RTA short, occupying about half tibial length, with distal end extended beyond tibia; lateral tibial apophysis large; cymbial furrow long, with distal end extended beyond cymbium, dorsal edge strongly concave; conductor moderately long; conductor dorsal apoph­ ysis long; conductor lamella strongly developed; embolus posterior in origin, long, linear; median apophysis spoonlike, with distal end not sharply pointed (figs. 183–185).

MATERIAL EXAMINED: CHINA: Hubei: Wudangshan, Jinding , August 1996, 1 male and 1 female paratype (J. Chen, HUW) ; Wudangshan, Jinding , September 24, 1997, 7 females (X. P. Wang, IZB) ; Wudangshan, Nanya to Jinding , September 24, 1997, 1 male and 16 females (X. P. Wang, IZB). Shaanxi: Taibaishan , Haoping , August 11, 1989, 1 male and 1 female, 1 male and 1 female, 1 male and 1 female, 1 male (X. P. Wang, AMNH and MCB) ; Taibaishan, Mingxinshi, Aug. 8, 1989, 1 female (X. P. Wang, IZB). Shanxi: Yongji , July 20, 1980, 1 female (M.S. Zhu, HUB, No­044) .

DISTRIBUTION: China (Hubei, Shanxi, Shaanxi).

Draconarius sinualis (Chen, Zhao and Wang) Figures 192–210

Coelotes sinualis Chen, Zhao and Wang, 1991: 10 , figs. 3, 4 (female holotype from Jinding , Wudangshan, Hubei, China, in HUW, examined); – Song et al., 1999: 378, figs. 224T, 224U.

DIAGNOSIS: Females can be distinguished by the strongly elongated, highly convoluted spermathecae, and the long, spiral copulatory ducts (fig. 192, 193), males by the long, slen­ der conductor and the slender conductor dorsal apophysis (figs. 198, 199).

DESCRIPTION: Eye pattern similar to D. wudangensis . Chilum divided (fig. 200). Chelicerae with three promarginal and three retromarginal teeth (fig. 201). Epigynum with epigynal teeth short, widely separated, situated anteral to atrium; atrium small; cop­ occupying half tibia, with distal end extended beyond tibia; lateral tibial apophysis large; cymbial furrow strongly elongated, with distal end extended beyond cymbium, dorsal edge strong concave; conductor long, slen­ der, slightly curved; conductor dorsal apophysis slender, long; conductor lamella strongly developed; embolus posterior in origin, extremely long, linear; median apophysis spoonlike, with distal end sharply pointed (figs. 198, 199).

Tarsal organ situated dorsally, away from distal tarsus, close to second distal trichobothrium (figs. 202–204). Apex of ALS with 2 major ampullate gland spigots (MAP), about 29 piriform gland spigots in female and 18 in male; PMS with 1 or 2 minor ampullate gland spigots in both sexes (not apparent), about 19 aciniform gland spigots in female and 12 in male, only 1 cylindrical gland spigot (CY) visible in female; PLS with about 20 aciniform gland spigots in both sexes and 2 cylindrical gland spigots in female (figs. 205–210).

MATERIAL EXAMINED: CHINA: Hubei: Wudangshan, Jinding , April 23, 1982, female holotype ( HUW) ; Wudangshan, Jinding , September 24, 1997, 3 females, 1 male and 2 females (X. P. Wang, AMNH) ; Wudangshan, Nanya to Jinding , September 24, 1997, 1 male and 1 female, 1 female, 11 females and 3 males (X. P. Wang, IZB) ; Wudangshan, Zhixiao to Nanya , September 23, 1997, 3 females and 7 males (X. P. Wang, MCB) ; Hongping, September 21, 1997, 2 males and 9 females (X. P. Wang, IZB). Shaanxi: Huxian, Cuihuashan , October 18, 1989, 1 male and 1 female (X. P. Wang, IZB) .

DISTRIBUTION: China (Hubei, Shaanxi).

ulatory ducts originated posteriorly, elongat­ ed, mesal to spermathecae; spermathecal heads small; spermathecae strongly elongat­ ed, highly convoluted, with indistinct stalks and bases (figs. 192–197).

Male patellar apophysis long; RTA short,

Figures

Figs. 181–185. Draconarius wudangensis (Chen and Zhao). 181. Female epigynum. 182. Female vulva. 183. Male palp, prolateral view. 184. Male palp, ventral view. 185. Male palp, retrolateral view.

Figs. 186–188. Draconarius wudangensis (Chen and Zhao), female. 186. Tarsus, showing position of tarsal organ (double arrows) and the distalmost trichobothrium (single arrow). 187. Trichobothrium. 188. Tarsal organ.

Figs. 189–191. Draconarius wudangensis (Chen and Zhao), female spinnerets, ventral view, left. 189. ALS. 190. PMS. 191. PLS. Abbreviations are spelled out on p. 6.

Figs. 192, 193. Draconarius sinualis (Chen, Zhao and Wang), female. 192. Epigynum. 193. Vulva.

Figs. 194–197. Draconarius sinualis (Chen, Zhao and Wang), female spermathecae. 194. Dorsal view. 195. Ventral view. 196. Dorsal view, enlarged, showing spermathecal pores situated on spermathecae. 197. Ventral view, enlarged, showing spermathecal head pores.

Figs. 198–201. Draconarius sinualis (Chen, Zhao and Wang). 198. Male palp, ventral view. 199. Male palp, retrolateral view. 200. Female, eyes, front view. 201. Female, chelicera, ventral view.

Figs. 202–204. Draconarius sinualis (Chen, Zhao and Wang), female. 202. Tarsus, showing position of tarsal organ (double arrows) and the distalmost trichobothrium (single arrow). 203. Tarsal organ. 204. Trichobothrium.