Leptocoelotes, WANG, 2002

WANG, XIN-PING, 2002, A Generic-Level Revision Of The Spider Subfamily Coelotinae (Araneae, Amaurobiidae), Bulletin of the American Museum of Natural History 2002 (269), pp. 1-150 : 105-109

LEPTOCOELOTES , NEW GENUS

TYPE SPECIES: Coelotes pseudoluniformis Zhang, Peng and Kim, 1998 .

ETYMOLOGY: The generic name is derived from its similarity to Coelotes and the weak epigynal teeth, and is masculine in gender.

DIAGNOSIS: Females can be easily recognized by the broad but weakly sclerotized epigynal teeth, a shallow atrium, and short copulatory ducts (figs. 296–299), and males by the complex conductor, the absence of a conductor dorsal apophysis, and the absence of median apophysis (figs. 302–304). Both sexes have four to five promarginal and retromarginal teeth (fig. 301).

DESCRIPTION: See description of type species (below).

DISTRIBUTION: China (map 11).

COMPOSITION: Two species are included.

1. Leptocoelotes edentulus ( Wang and Ono, 1998) : female holotype from Ilan, Taiwan, in NSMT, examined. NEW COMBINATION (transferred here from Coelotes ).

2. Leptocoelotes pseudoluniformis ( Zhang, Peng and Kim, 1998) : female holotype from Tiantong, Zhejiang, China, in HBI, examined. NEW COMBINATION (transferred here from Coelotes ).

Leptocoelotes pseudoluniformis (Zhang, Peng, and Kim) Figures 296–309

Coelotes pseudoluniformis Zhang, Peng and Kim, 1998: 293 , figs. 6, 7 (female holotype from Tiantong, Zhejiang, China, in HBI, examined).

DIAGNOSIS: Female of this species can be recognized from L. edentus by the laterally

situated hoods, the anterior origin of the copulatory ducts, and the relatively elongated spermathecae (figs. 296–299), and males by the tiny lateral tibial apophysis, the strongly bifid conductor, and the absence of both a conductor dorsal apophysis and a median apophysis (figs. 302–304).

DESCRIPTION: Total length about 5.00– 8.50. From front, anterior eye row slightly procurved, posterior row procurved; eye sizes and arrangements: AME smallest and slightly smaller than ALE, ALE subequal to PLE and PME; AME­AME slightly shorter than AME diameter, ALE­AME close together, subequal to ALE­PLE, PME­PLE subequal to PME­PME, about PME radius, AME­PME about PME radius. Clypeal height about 1.5–2 times AME diameter; chilum divided, elongated (fig. 300). Chelicerae with five promarginal and four to five retromarginal teeth (fig. 301). Labium with width slightly wider than length. Length of 1st leg patella + tibia longer than carapace length.

Tarsal organ situated away from distal end of tarsus, about same position as most distal trichobothrium (figs. 305, 306). Female spinnerets with apex of ALS with 2 major ampullate gland spigots (MAP), 19 piriform gland spigots; PMS with 1 or 2 minor ampullate gland spigots (mAP), 7 aciniform gland spigots, 2 cylindrical gland spigots (CY); PLS with about 12 aciniform gland spigots and 2 cylindrical spigots (figs. 307– 309).

Epigynum with epigynal teeth broad but weakly sclerotized; atrium broad but shallow, not apparent; copulatory ducts short, situated anterolaterad of spermathecae; spermathecal heads situated anterad of spermathecae; spermathecae slightly elongated, convoluted, widely separated, with indistinct bases and stalks (figs. 296–299). Male palpal patellar apophysis small, strongly curved dorsally; RTA small, with distal end extended beyond tibia; lateral tibial apophysis tiny; cymbial furrow moderately long; conductor complex; conductor dorsal apophysis absent; conductor lamella short; embolus basal in origin,

moderately long, linear; median apophysis absent (figs. 302–304).

MATERIAL EXAMINED: CHINA: Zhejiang: Tiantong, January 20, 1988, female holotype (Y. J. Zhang, HBI). Hunan: Changsha, Yuelushan, December 22, 1982, 2 males and 4 females (J. F. Wang, HBI).

2002 WANG: COELOTINAE 109

DISTRIBUTION: China (Zhejiang, Hunan).

Figures

Figs. 296, 297. Leptocoelotes pseudoluniformis (Zhang, Peng and Kim), female. 296. Epigynum.

Figs. 298–302. Leptocoelotes pseudoluniformis (Zhang, Peng and Kim). 298. Female, epigynum. 299. Female, vulva. 300. Carapace, front view. 301. Chelicera, ventral view. 302. Male palp, prolateral view.

Figs. 303, 304. Leptocoelotes pseudoluniformis (Zhang, Peng and Kim), male palp. 303. Ventral view. 304. Retrolateral view.

Figs. 305, 306. Leptocoelotes pseudoluniformis (Zhang, Peng and Kim), female. 305. Tarsus, showing position of trichobothria (single arrow) and tarsal organ (double arrows). 306. Trichobothrium and tarsal organ.

Figs. 307–309. Leptocoelotes pseudoluniformis (Zhang, Peng and Kim), female spinnerets, ventral view, left. 307. ALS. 308. PMS. 309. PLS. Abbreviations are spelled out on p. 6.