Platocoelotes, WANG, 2002

WANG, XIN-PING, 2002, A Generic-Level Revision Of The Spider Subfamily Coelotinae (Araneae, Amaurobiidae), Bulletin of the American Museum of Natural History 2002 (269), pp. 1-150 : 119-126

PLATOCOELOTES , NEW GENUS

TYPE SPECIES: Coelotes impletus Peng and Wang, 1997 .

ETYMOLOGY: This generic name is de­

364. Female, carapace, front view. 365. Female, chelicera, ventral view. 366. Male palp, prolateral view.

rived from its similarity to Coelotes and to its broad epigynum, and is masculine in gender.

DIAGNOSIS: Females can be easily recognized by the absence of epigynal teeth, the presence of an anteriorly situated epigynal cavity, the deep, posteriorly situated epigynal hoods, and the long, strongly convoluted spermathecae (figs. 336, 337), and males by the presence of two patellar apophyses (one strongly reduced in some specimens), the elongated cymbial furrow, the presence of a conductor posterior apophysis, a long embolus, and the absence of a median apophysis

(figs. 338–340). Both sexes have 3 promarginal and 2 retromarginal cheliceral teeth (fig. 342).

DESCRIPTION: See description of type species (below).

DISTRIBUTION: China (map 14).

COMPOSITION: Four species are included in this genus.

1. Platocoelotes impletus ( Peng and Wang, 1997) : female holotype, male and female paratypes from Zhangjiajie, Hunan, China, in HBI, examined. NEW COMBINATION (transferred here from Coelotes ).

2. Platocoelotes icohamatoides ( Peng and Wang, 1997) : female paratype from Naer Mt., Fenghuang, Hunan, China, in HBI, examined. NEW COMBINATION (transferred here from Coelotes ).

3. Platocoelotes icohamatus (Zhu and Wang, 1991) : female holotype, male and female paratypes from Sichuan, China, in NBUMS, not examined. NEW COMBINATION (transferred here from Coelotes ).

4. Platocoelotes lichuanensis (Chen and Zhao, 1998) : male holotype from Lichuan, Hubei, China, in HUW, examined. NEW COM­ BINATION (transferred here from Coelotes ).

Platocoelotes impletus (Peng and Wang) Figures 336–348

Coelotes impletus Peng and Wang, 1997: 328 , figs. 11–19 (female holotype, male and female paratypes from Zhangjiajie, Hunan, China, in HBI, examined). – Song et al., 1999: 375, figs. 220D, 220E, 222H, 223K.

DIAGNOSIS: Females are similar to those of

P. icohamatoides but can be distinguished by the different shape of epigynum and the long spermathecal stalks (figs. 336, 337). Males can be distinguished from those of P. lichuanensis only by the size of the conductor apex apophyses (figs. 338–340).

DESCRIPTION: Total length 5.00–12.0.

From front, anterior eye row slightly procurved, posterior row procurved; eye sizes and arrangements: AME slightly smaller than ALE, ALE subequal to PLE and PME; AME separated by roughly their radius, AME­ALE and ALE­PLE close together, PME­PLE

about 1.5 PME diameter, PME­PME separated by roughly their radius. Clypeal height about twice AME diameter or slightly less; chilum divided (fig. 341). Chelicerae with three promarginal and two retromarginal teeth (fig. 342). Labium about as wide as long. Length of female 1st leg patella + tibia longer than carapace length.

Tarsal organ not seen under SEM in examined specimens (figs. 343, 344). Further examination of other species also failed to locate tarsal organ (fig. 345). Female spinnerets examined: apex of ALS with 2 major ampullate gland spigots (MAP), 64 piriform gland spigots; PMS with at least 27 aciniform gland spigots, 2 cylindrical gland spig­ ots (CY), and 1 or 2 minor ampullate gland spigots (mAP); PLS with at least 44 aciniform gland spigots, cylindrical gland spigots not clear in examined specimen (figs. 346– 348).

Epigynum without epigynal teeth; hoods situated posteriorly near epigastric margin; atrium longitudinally elongated, shallow; epigynum with anteriorly situated, shallow cavity; copulatory ducts short; spermathecal heads small; spermathecae with stalks strongly elongated, highly convoluted; spermathecal bases small (figs. 336, 337). Male palp with two small patellar apophyses; RTA long, occupying almost tibial length, with distal end strongly extended beyond tibia; lateral tibial apophysis small; cymbial furrow long, with distal end strongly extended, slightly beyond cymbium; conductor broad, with long posteriorly extended apophysis and anteriorly extended finger­like apophyses; conductor dorsal apophysis broad; conductor lamella moderately developed; embolus posterior in origin and then curved prolaterally, long, linear; median apophysis absent (figs. 338–340).

MATERIAL EXAMINED: CHINA: Hunan: Zhangjiajie, August 18–20, 1988, 2 male and 2 female types (J. F. Wang and X. J. Peng, HBI).

DISTRIBUTION: China (Hunan).

Figures

Figs. 336, 337. Platocoelotes impletus (Peng and Wang), female. 336. Epigynum. 337. Vulva.

Figs. 338, 339. Platocoelotes impletus (Peng and Wang), male palp. 338. Ventral view. 339. Retrolateral view.

Figs. 340–342. Platocoelotes impletus (Peng and Wang), male palp. 340. Prolateral view. 341. Carapace, front view. 342. Chelicera, ventral view.