Spiricoelotes, WANG, 2002

WANG, XIN-PING, 2002, A Generic-Level Revision Of The Spider Subfamily Coelotinae (Araneae, Amaurobiidae), Bulletin of the American Museum of Natural History 2002 (269), pp. 1-150 : 129-133

SPIRICOELOTES NEW GENUS

TYPE SPECIES: Coelotes zonatus Peng and Wang, 1997 .

ETYMOLOGY: The generic name is derived from Coelotes and the spiral conductor, the strongly spiral, convoluted spermathecae, and is masculine in gender.

DIAGNOSIS: Females can be easily recognized by the the long, strongly convoluted spermathecae and the absence of epigynal teeth (figs. 360, 362), males by the strongly curved patellar apophysis, the elongated cymbial furrow, the absence of a conductor dorsal apophysis, and the slender, anteriorly extended, spiral conductor (figs. 366–368).

1. Spiricoelotes urumensis (Shimojana, 1989) : male holotype, male and female paratypes from Ishigaki City, Japan, in NSMT, not examined. NEW COMBINATION (transferred here from Coelotes ).

2. Spiricoelotes zonatus ( Peng and Wang, 1997) : male and female paratypes from Changsha, Hunan, China, in HBI, examined. NEW COMBINATION (transferred here from Coelotes ).

Both sexes have five teeth on both cheliceral margins (fig. 365).

DESCRIPTION: See description of type species (below).

DISTRIBUTION: China, Japan (map 16).

COMPOSITION: Two species are included in this genus.

Spiricoelotes zonatus (Peng and Wang)

Figures 360–374

Coelotes zonatus Peng and Wang, 1997: 331 , figs.

32–36 (male and female paratypes from Chang­

sha, Hunan, China, in HBI, examined). – Song

et al., 1999: 376, figs. 220H, 220I, 222I, 223L. Coelotes laoyingensis Chen and Zhao, 1997: 89 ,

figs. 5, 6 (female holotype and female paratype

from Laoying, Wudangshan, Hubei, China, in

HUW, examined). – Song et al., 1999: 388, figs.

226O, 226P, 227P, 229B. NEW SYNONYMY.

DIAGNOSIS: This species can be distinguished from S. urumensis by the strong spermathecae of females (fig. 361) and the relatively short conductor of males (fig. 367).

DESCRIPTION: Total length 6.00 –8.00. From front, anterior eye row slightly procurved, posterior row procurved; eye sizes and arrangements: AME smallest, slightly small­ er than ALE, ALE, PLE and PME subequal; AME separated by roughly three­fourths their diameter, AME­ALE and ALE­PLE close together, PME­PLE separated by roughly three­fourths their diameter, PME­ PME subequal to AME­AME or slightly longer. Clypeal height slightly longer than AME diameter; chilum divided (fig. 364). Chelicerae with five teeth on both margins (fig. 365). Labium wider than long. Length of female 1st leg patella + tibia longer than carapace length.

Tarsal organ situated anterior of most distal trichobothrium (figs. 369–371). Only female spinnerets examined: apex of ALS with 2 major ampullate gland spigots (MAP), about 23 piriform gland spigots; PMS with spigots with 1 or 2 minor ampullate gland spigots (mAP), about 20 aciniform gland spigots, 2 cylindrical gland spigots (CY); PLS with about 30 aciniform gland spigots, 2 cylindrical gland spigots (figs. 372–374).

Epigynum with epigynal teeth absent; atrium reduced to atrial slits; hoods well developed; copulatory ducts short; spermathecal heads small, apparent; spermathecae with stalks long, slender, spiral twice around copulatory ducts, then down close together; spermathecae bases slightly looped (figs. 360, 361). Male palpal patellar apophysis long, with distal end sharply curved dorsally; RTA long, with distal end broad, not extend anteriorly; lateral tibial apophysis present; cymbial furrow elongated, with distal end strongly extended beyond cymbium, dorsal edge strongly concave; conductor long, slen­ der, extended anteriorly; conductor dorsal apophysis absent; conductor lamella small; embolus posterior in origin, long, linear; median apophysis absent (figs. 366–368).

MATERIAL EXAMINED: CHINA: Hunan: Changsha , January 7, 1985, 2 male and 2 female paratypes (J. F. Wang, HBI) ; Changsha, Yuelushan , June 1995, 1 female (X. P. Wang, IZB). Sichuan: Chengdu, Bao­guang Monstery, May 21, 1989, 2 males (P. Beron,

COLL. DEELEMAN­REINHOLD). Jiangsu: Nanjing, Zijin (Purple) Mt., 350–450 m, October 9, 1988, 1 female (P. Beron, COLL. DEELEMAN­REINHOLD). Hubei: Wudangshan, Laoying, May 10, 1982, female holotype and female paratype of Coelotes laoyingensis (HUW) .

DISTRIBUTION: China (Hubei, Hunan, Jiangsu, Sichuan).

Figures

Figs. 360, 361. Spiricoelotes zonatus (Peng and Wang), female. 360. Epigynum. 361. Vulva.

Figs. 362–366. Spiricoelotes zonatus (Peng and Wang). 362. Female, epigynum. 363. Female, vulva.

Figs. 367, 368. Spiricoelotes zonatus (Peng and Wang), male palp. 367. Ventral view. 368. Retrolateral view.

Figs. 369–371. Spiricoelotes zonatus (Peng and Wang), female. 369. Tarsus, showing position of tarsal organ (double arrows) and distalmost trichobothrium (single arrow). 370. Trichobothrium. 371. Tarsal organ.