Ptomaphaminus ater Perreau, 2009

Schilthuizen, Menno, Perreau, Michel & Njunjic, Iva, 2018, A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae), ZooKeys 777, pp. 57-108 : 82-84

Ptomaphaminus ater Perreau, 2009 Figures 12e, f, 13e, f, 14d

Ptomaphaminus ater Perreau, 2009: 5, fig. 5; type from Gunung Kinabalu, Sabah, Borneo (in MHNG).

Ptomaphaginus ater : Merckx et al. 2015: extended data figs 2, 6.

Material.

(In addition to that listed in Perreau 2009): Sabah. Kinabalu Park, Paka Cave, 3080 m elev., 14-19.ix.2012 (leg. M. Schilthuizen, Crocker Range / Kinabalu Expedition, RMNH), 10 individuals (incl. RMNH.INS.555623-555624, RMNH.INS.1086288-1086290).

Description.

Length 1.75-2.00 mm. Large species, dark brown, winged. Eyes reduced, with 25 ommatidia. Pronotum ca. 1.5 times wider than long. Elytra approximately 1.2 times longer than wide. Female apex of the elytra with a sharp sutural angle expanded posteriorly (Figure 14d). Male protarsi approximately 0.6 times as wide as the apex of protibia. Male genital segment with long and apically expanded spiculum gastrale, widely dilated into a kind of paddle (Figure 13e). Aedeagus long, 3.5 times shorter than the body length, parallel-sided, straight, shortly narrowed near the apex in dorsal view (Figs 12f), ventrally bent on the last quarter of its length and with an apical hook clearly retroverted ventrally (Figure 12g). Stylet of the endophallus long and straight. Female genital segment with reduced gonocoxites. Spermiduct helical. Spermatheca long, bilobate, rounded at the apex, but with an apical sclerotised plate (Figure 13f).

DNA barcodes.

For two individuals, RMNH.INS.555623-555624, COI barcodes are available in BOLD, which form the BIN ACK0013.

Differential diagnosis.

Externally recognizable by its large size, dark colouration, and strongly cuneiform habitus. Apex of the female elytra spiniform. Spermatheca with a sclerotised plate at its apex. Aedeagus blunt-ended.

Habitat and distribution.

Known from high altitude on Gunung Kinabalu, above 3000 m. Some specimens were taken under a rocky overhang (Paka cave), others in Panar Laban (type locality) and Gunting Lagadan, without detail on collecting conditions.

Remarks.

The only species in Borneo with an apical sclerotised plate at the apex of the spermatheca (which occurs in several other species outside Borneo).