Wroughtonia jiangliae van Achterberg et Chen

Yan, Cheng-Jin, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2017, Review of the tribe Helconini Foerster s. s. from China, with the description of 18 new species, Zootaxa 4291 (3), pp. 401-457 : 436-438

Wroughtonia jiangliae van Achterberg et Chen , sp. nov.

( Fig. 18 )

Description. Female. Length of body (excluding ovipositor sheath) 9.1 mm, of setose part of ovipositor sheath 8.4 mm, of fore wing 7.2 mm.

Head. Antennal segments 38, length of third segment 1.1 times fourth segment; length of third, fourth and penultimate segments 3.4, 2.8 and 1.6 times their width, respectively; length of maxillary palp 1.4 times height of head; head 0.8 times as long as wide in anterior view; length of eye 1.5 times temple in dorsal view; length of malar space 1.1 times basal width of mandible and 0.5 times maximum width of eye; malar suture moderately impressed, rugose; POL:OD:OOL = 9:5:16; vertex sparsely punctate and shiny, but with larger punctures near eye ( Fig. 18 I); temple coarsely and rather spaced punctate, with some rugae ventrally; occipital carina complete, gradually curved ventrally and meeting hypostomal carina far above base of mandible, occipital flange large, lamelliform and upcurved ( Fig. 18 I); frontal protuberance wide triangular, largely smooth laterally and pointing dorsally, slightly concave posteriorly, in posterior view narrow triangular and depressed basally ( Fig. 18 J), frontal depression deep anteriorly, and rather so posteriorly, with coarse sublateral rugae, medially coarsely vermiculate-rugose; face coarsely vermiculate-rugose; clypeus weakly convex, dorsally densely rugose and ventrally depressed and mainly smooth; epistomal suture deep.

Mesosoma. Length 1.8 times its height; pronope obsolescent ( Fig. 18 D); propleuron strongly convex and rugulose; side of pronotum coarsely vermiculate-rugose, but finely punctate dorsally; mesoscutal lobes largely smooth except some fine punctulation, vermiculate-rugose laterally, near notauli and posteriorly; notauli wide and deep, rugose, posteriorly with a median carina ( Fig. 18 D); scutellar sulcus wide and with four carinae ( Fig.18 D); scutellum sparsely punctulate, nearly flat and without lateral carinae; precoxal sulcus narrow, with some spaced coarse punctures; metanotum with a complete median carina, posteriorly triangularly protruding; propodeum narrowly smooth basally except some punctures, behind distinct transverse and curved carina mainly coarsely foveolate-rugose, with irregular areola and with short median carina ( Fig. 18 D).

Wings ( Fig. 18 A). Fore wing: about 4 times as long as wide. 1-M slightly curved; pterostigma 3.2 times as long as wide, elliptical; r:3-SR:SR1 = 9:10:59; 2-SR:3-SR:r-m = 17:10:12; 1-M:m-cu = 19:10; SR1 nearly straight; cu-a strongly inclivous and postfurcal; 1-CU1:2-CU1 = 1:8; r-m distinctly inclivous. Hind wing: 1-M:1r-m = 5:11; cu-a slightly curved posteriorly; marginal cell distinctly widened.

Legs. Length of fore tarsus 1.2 times fore tibia; length of femur, tibia and basitarsus of hind leg 3.2 (including protuberance), 8.5 and 6.9 times their width, respectively; length of outer and inner hind tibia spur 0.25 and 0.20 times basitarsus, respectively and outer spur widened; hind femur robust, ventrally rugose and with a narrow flange-like protuberance ( Fig. 18 F); hind coxa stout, dorsally largely smooth, except for fine spaced punctation.

Metasoma. First tergite distinctly widened posteriorly, apically 3.5 times wider than basally, coarsely foveolate-punctate and its dorsal carinae nearly complete; length of first tergite equal to its apical width; second tergite coarsely foveolate-punctate ( Fig. 18 E), except posteriorly; second suture narrow and punctate; third tergite moderately foveolate-punctate but medially and posteriorly smooth ( Fig. 18 E); ovipositor sheath 2.1 times as long as metasoma, 2.2 times as long as hind tibia and 1.16 times as long as fore wing; ovipositor sheath with fine rather adpressed brownish setae, becoming paler apically and with some long apical setae ( Fig. 18 G).

Colour. Black; palpi (including basal segments) pale yellow; antenna dark brown or black; all tarsi more or less ivory; coxae black; tegulum, hind femur (except narrowly yellowish brown base) and tibia (except pale base), trochantelli and hind trochantellus dark brown; remainder of legs brownish yellow; metasoma dark brown or brown ventrally and black dorsally, third tergite apically reddish brown; ovipositor sheath and setae dark brown with paler setae apically; pterostigma and most veins dark brown; wing membrane faintly fumose.

Variation. Length of body 9.1-10.2 mm and of fore wing 7.2-7.5 mm; antenna of ♀ with 37(1) or 38(2) segments; length of ovipositor sheath 0.93-1.16 times fore wing; basal half of hind femur largely dark brown or brown.

Material examined. Holotype, ♀, NW. China, Shaanxi prov., Qinling Mts, Baolongyu , c. 1000 m, 25.V.-10.VI.2015, 34°03’N 108°09’E, Malaise trap, Jiangli Tan ( NWUX) . Paratype: 1♀ NW. China: Shaanxi prov., Qinling Mts, Foping, behind Biological Station , Malaise trap, 33˚39'29"N 107˚48'25"E, 29.V.-19.VI.2016, c 1710 m, JL. Tan & C. v. Achterberg ( RMNH) .

Comparative diagnosis. The entirely dark antenna and the black first metasomal tergite immediately separates W. jiangliae from the other Chinese species with third tergite sculptured and fourth tergite more or less retracted (“ Spasskia ”). It shares this with W. sigalphoides ( Belokobylskij, 1989) comb. nov. from Far East Russia. Differs by having vertex and temple sparsely punctate (entirely coarsely rugose-areolate in W. sigalphoides ), mesoscutum medially remotely finely punctate (mainly rugose-punctate); pterostigma elliptical (triangular), frontal protuberance triangular dorsally (distinctly bifurcate dorsally), hind femur behind small flange distinctly concave (slightly concave), precoxal sulcus narrow, with some spaced coarse punctures (coarsely and widely rugosereticulate), basal half of hind femur mainly dark brown (yellowish brown) and pterostigma elliptical (triangular).

Host. Unknown.

Etymology. It is named after its collector, Dr Jiangli Tan (NWUX) for her efforts to sample the Hymenoptera fauna of China, especially of Shaanxi province.

Figures

FIGURE 18. Wroughtonia jiangliae sp. nov., ♀, holotype. A, fore wing; B, hind wing; C, mesosoma, lateral aspect; D, mesosoma, dorsal aspect; E, metasoma, dorsal aspect; F, hind leg, lateral aspect; G, apical part of ovipositor sheath, lateral aspect; H, habitus, lateral aspect; I, head, lateral aspect; J, head, dorsal aspect; K, face and clypeus, anterior aspect.